A review of 21st century studies on lizards (Reptilia: Squamata: Sauria) in northeastern India with an updated regional checklist
DOI:
https://doi.org/10.11609/jott.8746.17.10.27720-27733Keywords:
Diversity, ecology, myths, range extension, rediscovery, saurian research, species description, systematicsAbstract
The current study reviews saurian research undertaken in the 21st century in northeastern India. The scope of this review encompasses new species descriptions, range extensions, diversity assessments, systematic revisions, and species rediscoveries. In addition, it incorporates miscellaneous contributions, particularly those addressing myths, ecological aspects, conservation threats, and human-wildlife interactions documented from different states of the region. Based on published works from northeastern India, an updated checklist of the region’s lizard fauna is presented. The checklist comprises 84 species representing six families and 19 genera, with their conservation status assessed according to the International Union for Conservation of Nature (IUCN) Red List of Threatened Species.
References
Agarwal, I., S. Biswas, A.M. Bauer, E. Greenbaum, T.R. Jackman, A.D. Silva & S. Batuwita (2017). Cryptic species, taxonomic inflation, or a bit of both? New species phenomenon in Sri Lanka as suggested by a phylogeny of dwarf geckos (Reptilia, Squamata, Gekkonidae, Cnemaspis). Systematics and Biodiversity 15(5): 427–439. https://doi.org/10.1080/14772000.2017.1282553
Agarwal, I., S. Mahony, V.B. Giri, R. Chaitanya & A.M. Bauer (2018a). Two new species of bent-toed geckos, Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from northeastern India with comments on name-bearing types from the region. Zootaxa 4420(3): 334–356. https://doi.org/10.11646/zootaxa.4420.3.2
Agarwal, I., S. Mahony, V.B. Giri, R. Chaitanya & A.M. Bauer (2018b). Six new Cyrtodactylus (Squamata: Gekkonidae) from northeastern India. Zootaxa 4524(5): 501–535. https://doi.org/10.11646/zootaxa.4524.5.1
Agarwal, I., V.K. Mistry & R. Athreya (2010). A preliminary checklist of the reptiles of Eaglenest Wildlife Sanctuary, West Kameng District, Arunachal Pradesh, India. Russian Journal of Herpetology 17(2): 81–93.
Ahmed, M.F., A. Das & S.K. Dutta (2009). Amphibians and Reptiles of Northeastern India: A Photographic Guide. Aaranyak, Guwahati, Assam, India, 169 pp.
Alcock, A. (1895). On a new species of flying lizard from Assam. The Journal of the Asiatic Society of Bengal 64(1): 14–15.
Annandale, N. (1905). Notes on some Oriental geckos in the Indian Museum, Calcutta, with descriptions of new forms. The Annals and Magazine of Natural History, including Zoology, Botany and Geology 7(15): 26–32.
Annandale, N. (1908). Description of a new species of lizard of the genus Salea from Assam. Records of the Zoological Survey of India 2(1): 37–38. https://doi.org/10.26515/rzsi/v2/i1/1908/163304
Annandale, N. (1912). Zoological results of the Abor Expedition, 1911–1912. Records of the Indian Museum 8(1): 7–59 [Reptilia: 37–59]. Supplement: Records of the Indian Museum 8(4): 357–358 (1914).
Annandale, N. (1913). The Indian geckos of the genus Gymnodactylus. Records of the Zoological Survey of India 9(4): 99–109. https://doi.org/10.26515/rzsi/v9/i4/1913/163665
Bharali, M., K. Thaosen, M. Vabeiryureilai, H.T. Lalremsanga, J. Purkayastha, R. Bhattacharjee, M. Das, S.C. Bohra, A.K. Hazarika (in press). A new species of Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae) from the montane forests of Dima Hasao District, Assam, India. Journal of Asia-Pacific Biodiversity. Available online 18 July 2025. https://doi.org/10.1016/j.japb.2025.06.008
Bhupathy, S., B. Chettri & A.M. Bauer (2009). Rediscovery and revalidation of Takydromus sikkimensis (Günther, 1888) (Squamata: Lacertidae) from Sikkim, India. Journal of Herpetology 43(2): 267–274. https://doi.org/10.1670/08-136R1.1
Bleeker, P. (1860). Reptilien van Agam. Natuurkundig Tijdschrift voor Nederlandsch Indie 20: 325–329.
Blyth, E. (1854). Notices and descriptions of various reptiles, new or little known. The Journal of the Asiatic Society of Bengal 22: 639–655.
Bohra, S.C., H.T. Zonunsanga, M. Das, J. Purkayastha, L. Biakzuala & H.T. Lalremsanga (2022). Morphological and molecular phylogenetic data reveal another new species of bent-toed gecko (Cyrtodactylus Gray: Squamata: Gekkonidae) from Mizoram, India. Journal of Natural History 56(41–44): 1585–1608. https://doi.org/10.1080/00222933.2022.2119178
Bohra, S.C., C. Ranglong, G. Thongni, B. Wankhar, C.M. Umlong, M. Das, H. Warjri & J. Purkayastha (2025). Leaping from Mizoram to Meghalaya: first documentation of Calotes zolaiking Giri, Chaitanya, Mahony, Lalronunga, Lalrinchhana, Das, Sarkar, Karanth & Deepak, 2019 (Sauria: Agamidae) outside its type locality in Mizoram, India. Zootaxa 5642(4): 395–400. https://doi.org/10.11646/zootaxa.5642.4.7
Boruah, B., S. Narayanan, N.A. Aravind, S. Lalronunga, V. Deepak & A. Das (2024). Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India. Vertebrate Zoology 74: 453–486. https://doi.org/10.3897/vz.74.e124752
Boruah, B., S. Narayanan, V. Deepak & A. Das (2022). Morphological and molecular differences in two closely related Calotes Cuvier, 1817 (Squamata: Agamidae: Draconinae) with the first record of Calotes medogensis Zhao & Li, 1984 from India. Zootaxa 5219(5): 433–455. https://doi.org/10.11646/zootaxa.5219.5.3
Boulenger, G.A. (1885). Catalogue of the Lizards in the British Museum (Natural History). Printed by order of the Trustees, London, 428 pp.
Boulenger, G.A. (1887). An account of the scincoid lizards collected in Burma, for the Genoa Civic Museum, by Messrs. G.B. Comotto & L. Fea. Annali del Museo Civico di Storia Naturale di Genova 4(2): 618–624.
Boulenger, G.A. (1890). The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, 541 pp.
Boulenger, G.A. (1903). Descriptions of new lizards in the collection of the British Museum. Annals and Magazine of Natural History 12(7): 429–435.
Boulenger, G.A. (1917). A revision of the lizards of the genus Tachydromus. Journal of the Asiatic Society of Bengal 13(4): 207–235.
Chandramouli, S.R., R.S. Naveen, S. Sureshmarimuthu, S. Babu, P.V. Karunakaran & H.N. Kumara (2021). Catalogue of herpetological specimens from Meghalaya, India at the Sálim Ali Centre for Ornithology and Natural History. Journal of Threatened Taxa 13(11): 19603–19610. https://doi.org/10.11609/jott.7318.13.11.19603-19610
Che, J., K. Jiang, F. Yan & Y. Zhang (2020). Amphibians and Reptiles in Tibet—Diversity and Evolution. Chinese Academy of Sciences, Beijing, 803 pp.
Chettri, B., S. Bhupathy & B.K. Acharya (2009). Distribution pattern of reptiles along an eastern Himalayan elevation gradient, India. Acta Oecologica 36(1): 16–22. https://doi.org/10.1016/j.actao.2009.09.004
Choudhury, A.S. & P. Choudhury (2019). Cruelty to Varanus species of the Barak Valley, Assam, India. Biawak 13(1): 50–53.
Das, A. & I. Das (2007). Rediscovery of Mictopholis austeniana (Annandale, 1908) (Squamata: Agamidae). Current Herpetology 26(1): 45–47.
Das, A. & M.F. Ahmed (2007). Range extension of Assamese Day Gekko Cnemaspis assamensis Das and Sengupta (Sauria: Gekkonidae). Zoos’ Print Journal 22(6): 2720. https://doi.org/10.11609/JoTT.ZPJ.1513a.2720
Das, A., M.F. Ahmed, B.P. Lahkar & P. Sharma (2007). A preliminary report of reptilian mortality on road due to vehicular movement near Kaziranga National Park, Assam, India. Zoos’ Print Journal 22(7): 2742–2744. https://doi.org/10.11609/JoTT.ZPJ.1541.2742-4
Das, A., U. Saikia, B.H.C.K. Murthy, S. Dey & S.K. Dutta (2009). A herpetofaunal inventory of Barail Wildlife Sanctuary and adjacent regions, Assam, north-eastern India. Hamadryad 34(1): 117–134.
Das, I. & S. Sengupta (2000). A new species of Cnemaspis (Sauria: Gekkonidae) from Assam, northeastern India. Journal of South Asian Natural History 5(1): 17–24.
Das, M., J. Purkayastha, A.M. Bauer & S. Sengupta (2011a). Hemidactylus flaviviridis Rüppell, 1835 (Sauria: Gekkonidae), an invasive gecko in Assam. North-Western Journal of Zoology 7(1): 98–104.
Das, M., R.K. Brahma & J. Purkayastha (2011b). More in our mind than in their mouth? A preliminary inspection inside the oral cavity of two house geckos: Hemidactylus frenatus Schlegel, 1836 and Hemidactylus aquilonius McMahan & Zug, 2007. Herpetology Notes 4: 303–306.
Datta-Roy, A., I. Das, A.M. Bauer, R.K.L. Tron & P. Karanth (2013). Lizard wears shades. A spectacled Sphenomorphus (Squamata: Scincidae) from the sacred forests of Mawphlang, Meghalaya, northeastern India. Zootaxa 3701(2): 257–276. https://doi.org/10.11646/zootaxa.3701.2.7
Daudin, F.M. (1802). Histoire naturelle, générale et particulière des reptiles, ouvrage faisant suite à l’histoire naturelle, générale et particulière composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, Vol. 1. F. Dufart, Paris, 452 pp.
Decemson, H., R.R. Sinate, L. Biakzuala, V. Mathipi, P. Lalnuntluanga, J. Malsawma & H.T. Lalremsanga (2023). Confirmation of Irawadi Forest Lizard, Calotes irawadi Zug, Brown, Schulte and Vindum 2006 (Squamata: Agamidae), from Manipur, northeastern India. Reptiles & Amphibians 30(1): e17043. https://doi.org/10.17161/randa.v30i1.17043
Decemson, H.T., M. Vabeiryureilai, L. Lalbiakzuala & H.T. Lalremsanga (2021). Confirmation on the occurrence of Calotes geissleri Wagner, Ihlow, Hartman, Flecks, Schmitz and Böhme, 2021 (Sauria: Agamidae) in Chandel, Manipur, India with comments on its phylogenetic position. Journal of Animal Diversity 3(4): 14–19. http://doi.org/10.52547/JAD.2021.3.4.2
Deepak, C.K., D. Kaushik & C. Kailash (2022). First record of Hemiphyllodactylus Bleeker, 1860 (Squamata: Sauria: Gekkonidae) from the Eastern Himalayas. Russian Journal of Herpetology 29(6): 367–372. https://doi.org/10.30906/1026-2296-2022-29-6-367-372
Duméril, A.M.C. & G. Bibron (1836). Erpétologie Générale ou Histoire Naturelle Complète des Reptiles. Vol. 3. Librairie Encyclopédique Roret, Paris, 528 pp.
Duméril, A.M.C. & G. Bibron (1837). Erpétologie Générale ou Histoire Naturelle Complète des Reptiles. Vol. 4. Librairie Encyclopédique Roret, Paris, 570 pp.
Giri, V.B., R. Chaitanya, S. Mahony, S. Lalrounga, C. Lalrinchhana, A. Das, V. Sarkar, P. Karanth & V. Deepak (2019). On the systematic status of the genus Oriocalotes Günther, 1864 (Squamata: Agamidae: Draconinae) with the description of a new species from Mizoram State, northeastern India. Zootaxa 4638(4): 451–484. https://doi.org/10.11646/zootaxa.4638.4.1
Gouda, S., H. Decemson, F. Malsawmdawngliana, L. Biakzuala & H.T. Lalremsanga (2023). Threat assessment and conservation challenges for the herpetofaunal diversity of Dampa Tiger Reserve, Mizoram, India. Journal of Threatened Taxa 15(10): 24016–24031. https://doi.org/10.11609/jott.8590.15.10.24016-24031
Gowande, G.G., H.S. Bhosale, P.U. Phansalkar, M. Sawant & Z.A. Mirza (2021). On the systematics and the phylogenetic position of the poorly known, montane dragon-lizard species Pseudocalotes austeniana (Annandale, 1908) (Squamata: Agamidae: Draconinae). Evolutionary Systematics 5(1): 141–150. https://doi.org/10.3897/evolsyst.5.67137
Gray, J.E. (1827). A synopsis of the genera of saurian reptiles in which some new genera are indicated, and the others reviewed by actual examination. Philosophical Magazine, London 2(2): 54–58.
Gray, J.E. (1845). Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of the British Museum/Edward Newman, London, xxvii + 289 pp.
Gray, J.E. (1846). Descriptions of some new species of Indian Lizards. Annals and Magazine of Natural History 18(1): 429–430.
Gray, J.E. (1853). Descriptions of some undescribed species of reptiles collected by Dr. Joseph Hooker in the Khassia Mountains, east Bengal, and Sikkim Himalaya. Annals and Magazine of Natural History 12(2): 386–392.
Grismer, L.L., P.L. Wood Jr., N.A. Poyarkov, M.D. Le, F. Kraus, I. Agarwal, P.M. Oliver, S.N. Nguyen, T.Q. Nguyen, S. Karunarathna, L.J. Welton, B.L. Stuart, V.Q. Luu, A.M. Bauer, K.A. O’Connell, E.S.H. Quah, K.O. Chan, T. Ziegler, H. Ngo, R.A. Nazarov, A. Aowphol, S. Chomdej, C. Suwannapoom, C.D. Siler, S. Anuar, N.V. Tri & J.L. Grismer (2021). Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154. https://doi.org/10.3897/vz.71.e59307
Grismer, L.L., M.K. Thura, M.S. Grismer, R.M. Brown & B.L. Stuart (2018). Geographically structured genetic variation in Ptychozoon lionotum (Squamata: Gekkonidae) and a new species from an isolated volcano in Myanmar. Zootaxa 4514(2): 202–214. https://doi.org/10.11646/zootaxa.4514.2.4
Günther, A. (1864). The Reptiles of British India. Ray Society, London, xxvii + 452 pp.
Günther, A. (1870). Descriptions of a new Indian lizard of the genus Calotes. Proceedings of the Zoological Society of London 1870: 778–779.
Günther, A. (1888). On a collection of reptiles from China. Journal of Natural History 1(3): 165–172.
Harit, D.N. (2018). Report on reptile mortality due to vehicular traffic on Indo-Myanmar border at Champhai District of Mizoram, north east India. Environment Conservation Journal 19(3): 133–138.
Islam, M. & P.K. Saikia (2013). Inventory and natural history of lizards in Jeypore Reserve Forest, Assam. Reptile Rap 16(1): 17–26.
Jerdon, T.C. (1870). Notes on Indian herpetology. Proceedings of the Asiatic Society of Bengal 1870: 66–85.
Jha, A. & K. Thapa (2002). Reptiles and Amphibians of Sikkim. Mrs. Shila Jha, Chhattisgarh, 100 pp.
Kamei, R.G. & S. Mahony (2021). A new species of Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus Gray, 1827) from the Garo Hills, Meghalaya State, north-east India, and discussion of morphological variation for C. urbanus. Herpetological Journal 31(3): 177–196. https://doi.org/10.33256/31.3.177196
Kunte, K. & U. Manthey (2009). Wiederentdeckung von Japalura sagittifera (Sauria: Agamidae) in Arunachal Pradesh, Ost-Himalaya: Ein Erstnachweis für die indische Herpetofauna. Sauria 31: 49–55.
Lalremsanga, H.T., Z. Colney, M. Vabeiryureilai, F. Malsawmdawngliana, S.C. Bohra, L. Biakzuala, L. Muansanga, M. Das & J. Purkayastha (2023a). It’s all in the name: Another new Cyrtodactylus Gray (Squamata: Gekkonidae) from northern Mizoram, north-east India. Zootaxa 5369(4): 553–575. https://doi.org/10.11646/zootaxa.5369.4.5
Lalremsanga, H.T., H. Chinliansiama, S.C. Bohra, L. Biakzuala, M. Vabeiryureilai, L. Muansanga, F. Malsawmdawngliana, G.Z. Hmar, H.T. Decemson, V. Siammawii, M. Das & J. Purkayastha (2022b). A new bent-toed gecko (Cyrtodactylus Gray: Squamata: Gekkonidae) from the state of Mizoram, India. Zootaxa 5093(4): 465–482. https://doi.org/10.11646/zootaxa.5093.4.5
Lalremsanga, H.T., H. Decemson, M. Vabeiryureilai, F. Malsawmdawngliana, V. Lalhlimpuia, L. Muansanga & L. Biakzuala (2022a). Phylogenetic position of Tropidophorus assamensis Annandale, 1912 with updated morphological data and distributional records. Herpetological Journal 32(1): 1–4. https://doi.org/10.33256/32.1.14
Lalremsanga, H.T., L. Khawlhring & Lalrotluanga (2010). Three additional lizard (Squamata: Sauria) records for Mizoram, India. Journal of Threatened Taxa 2(2): 718–720. https://doi.org/10.11609/JoTT.o2246.718-20
Lalremsanga, H.T., L. Muansanga, M. Vabeiryureilai & Z.A. Mirza (2023b). A new species of Parachute Gecko of the subgenus Ptychozoon (Sauria: Gekkonidae: Gekko) from the Indo-Burma region. Salamandra 59(2): 125–135.
Lalrinchhana, C. & G.S. Solanki (2015). Lizard (Reptilia: Sauria) diversity of Dampa Tiger Reserve, Mizoram, India. Science Vision 15(1): 19–28.
Laurenti, J.N. (1768). Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austracorum, quod authoritate et consensus. Joan. Thomae, Vienna, 217 pp.
Linnaeus, C. (1758). Systema naturae, per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiæ, 824 pp.
Mahananda, P., S.N. Jelil, S.C. Bohra, N. Mahanta, R.B. Saikia & J. Purkayastha (2023). Terrestrial vertebrate and butterfly diversity of Garbhanga Landscape, Assam, India. Journal of Threatened Taxa 15(4): 23029–23046. https://doi.org/10.11609/jott.8334.15.4.23029-23046
Mahony, S. & R.G. Kamei (2022). A new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Manipur State, northeastern India, with a critical review highlighting extensive errors in literature covering bent-toed geckos of the Indo-Burma region. Journal of Natural History 55(39–40): 2445–2480. https://doi.org/10.1080/00222933.2021.1994667
Mahony, S. (2009). A new species of Japalura (Reptilia: Agamidae) from northeastern Indiawith a discussion of the similar species Japalura sagittifera Smith, 1940 and Japalura planidorsata Jerdon, 1870. Zootaxa 2212(1): 41–61. https://doi.org/10.11646/zootaxa.2212.1.2
Majumder, J. & B.K. Agarwala (2015). Calotes emma Gray, 1845 (Squamata: Agamidae): range extension and new addition to the reptilian fauna of Tripura, northeastern India. Check List 11(2): 1562. https://doi.org/10.15560/11.2.1562
Majumder, J., P.P. Bhattacharjee, K. Majumdar, C. Debnath & B.K. Agarwala (2012). Documentation of herpetofaunal species richness in Tripura, northeastern India. NeBio: An International Journal of Environment and Biodiversity 3(1): 60–70.
Malsawmdawngliana, B. Boruah, N.G. Patel, S. Lalronunga, I. Zosangliana, K. Lalhmangaiha & A. Das (2022). An updated checklist of reptiles from Dampa Tiger Reserve, Mizoram, India, with sixteen new distribution records. Journal of Threatened Taxa 14(10): 21946–21960. https://doi.org/10.11609/jott.8004.14.10.21946-21960
Mathew, R. (1995). State Fauna Series 4: Fauna of Meghalaya, Part 1. Zoological Survey of India, Kolkata: 379–454.
McMahan, C.D. & G.R. Zug (2007). Burmese Hemidactylus (Reptilia: Squamata: Gekkonidae): geographic variation in the morphology of Hemidactylus bowringii in Myanmar and Yunnan. Proceedings of the California Academy of Sciences 58(23–30): 485–509.
Mirza, Z.A., A.M. Bragin, H. Bhosale, G.G. Gowande, H. Patel & N.A. Poyarkov (2022a). A new ancient lineage of ablepharine skinks (Sauria: Scincidae) from eastern Himalayas with notes on origin and systematics of the group. PeerJ 10(1): e12800. https://doi.org/10.7717/peerj.12800
Mirza, Z.A., H. Bhosale, F. Ansari, P. Phansalkar, M. Sawant, G. Gowande & H. Patel (2021). A new species of geckos of the genus Cyrtodactylus Gray, 1827 from Arunachal Pradesh, India. Evolutionary Systematics 5(1): 13–23. https://doi.org/10.3897/evolsyst.5.61667
Mirza, Z.A., H.S. Bhosale, T. Thackeray, P. Phansalkar, M. Sawant, G.G. Gowande & H. Patel (2022b). A new species of bent-toed geckos of the genus Cyrtodactylus Gray, 1827 from western Arunachal Pradesh, India. Herpetozoa 35(1): 65–76. https://doi.org/10.3897/herpetozoa.35.e80610
Mirza, Z.A., G. Gowande, T. Thackeray, H. Bhosale, M. Sawant, P. Phansalkar & H. Patel (2024). A new montane-dwelling species of Japalura Gray, 1853 (Squamata: Agamidae) from Arunachal Pradesh, India. Taprobanica 13(1): 1–8. https://doi.org/10.47605/tapro.v13i1.317
Muansanga, L., H. Decemson, L. Biakzuala, G.Z. Hmar, H.T. Lalremsanga, M. Das & J. Purkayastha (2020). First record of the Jampui Bent-toed Gecko, Cyrtodactylus montanus Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 (Squamata: Gekkonidae), from Mizoram, India. Reptiles & Amphibians 27(2): 267–268. https://doi.org/10.17161/randa.v27i2.14325
Patel, H., T. Thackeray, C. Sheth, A. Khandekar & I. Agarwal (2024). A new small-sized Calotes Cuvier (Squamata: Agamidae: Draconinae) from the Subansiri River Basin, Arunachal Pradesh, India. Zootaxa 5523(2): 151–170. https://doi.org/10.11646/zootaxa.5523.2.1
Pawar, S.S., G.S. Rawat & B.C. Choudhury (2004). Recovery of frog and lizard communities following primary habitat alteration in Mizoram, northeastern India. BMC Ecology 4(1): 1–18. https://doi.org/10.1186/1472-6785-4-10
Purkayastha, J. (2013). An Amateur’s Guide to Reptiles of Assam. Eastern Book House Publishers (India), Guwahati, 144 pp.
Purkayastha, J. (2018). Urban biodiversity: an insight into the terrestrial vertebrate diversity of Guwahati, India. Journal of Threatened Taxa 10(10): 12299–12316. https://doi.org/10.11609/jott.3721.10.10.12299-12316
Purkayastha, J., S.C. Bohra & M. Das (2020d). First record of the Urban Bent-toed Gecko, Cyrtodactylus urbanus Purkayastha, Das, Bohra, Bauer, and Agarwal, 2018 (Squamata: Gekkonidae), from Meghalaya, India. Reptiles & Amphibians 27(3): 512–513. https://doi.org/10.17161/randa.v27i3.14901
Purkayastha, J., H.T. Lalremsanga, B. Litho, Y.S. Rathee, S.C. Bohra, M. Vabeiryureilai, L. Biakzuala & L. Muansanga (2022). Two new Cyrtodactylus (Squamata: Gekkonidae) from northeastern India. European Journal of Taxonomy 794: 111–139. https://doi.org/10.5852/ejt.2022.794.1659
Purkayastha, J., H.T. Lalremsanga, S.C. Bohra, L. Biakzuala, H.T. Decemson, L. Muansanga, M. Vabeiryureilai, S. Chauhan & Y.S. Rathee (2021). Four new Bent-toed Geckos Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from northeastern India. Zootaxa 4980(3): 451–489. https://doi.org/10.11646/zootaxa.4980.3.2
Purkayastha, J., M. Das & S. Sengupta (2011). Urban herpetofauna: a case study in Guwahati City of Assam, India. Herpetology Notes 4(1): 195–202.
Purkayastha, J., M. Das, A.M. Bauer, S. Sengupta & S.K. Dutta (2010). Notes on the Hemidactylus bowringii complex (Reptilia: Gekkonidae) in India, and a change to the national herpetofaunal list. Hamadryad 35(1): 20–27.
Purkayastha, J., M. Das, S.C. Bohra, A.M. Bauer & I. Agarwal (2020a). Another new Cyrtodactylus (Squamata: Gekkonidae) from Guwahati, Assam, India. Zootaxa 4732(3): 375–392. https://doi.org/10.11646/zootaxa.4732.3.2
Purkayastha, J., N. Khan & S. Roychoudhury (2020b). A preliminary checklist of herpetofauna occurring in Rowa Wildlife Sanctuary, Tripura, India, pp. 225–233. In: Förstner, U. & W.H. Rulkens (eds.). Environmental Science and Engineering. Springer International Publishing. https://doi.org/10.1007/978-3-030-32463-6_10
Purkayastha, J., S. Roychoudhury, B.B. Biswa, M. Das & S. Sengupta (2020c). Herpetofaunal Diversity and conservation status in Amchang Wildlife Sanctuary of Assam, India, pp. 217–223. In: Roy, N., S. Roychoudhury, S. Nautiyal, S. Agarwal & S. Baksi (eds.). Socio-economic and Ecobiological Dimensions in Resource Use and Conservation: Strategies for Sustainability. Springer, Cham.
Rai, A., R. Mothey & B. Chettri (2023). Sexual dimorphism and reproductive biology of a Sikkim Endemic Grass Lizard Takydromus sikkimensis (Squamata: Lacertidae) in the eastern Himalaya, India. Herpetological Conservation and Biology 18: 508–519.
Ranade, S. & J. Purkayastha (2020). Cannibalism in the Kachin Smooth Gecko, Hemidactylus aquilonius Zug & McMahan, 2007 (Squamata: Gekkonidae), from Assam, India. Reptiles & Amphibians 27(3): 514. https://doi.org/10.17161/randa.v27i3.14902
Rüppell, E. (1835). Neue Wirbelthiere zu der Fauna von Abyssinien gehörig, entdeckt und beschrieben. Amphibien. S. Schmerber, Frankfurt am Main, 18 pp.
Sauvage, H.E. (1879). Notice sur quelques reptiles nouveaux ou peu connus de la Nouvelle-Guinée. Bulletin des Sciences par la Société Philomatique de Paris, Série 7(3): 47–61.
Sayyed, A., M. Das, A.A.T. Amarasinghe, R. Bhattacharjee & J. Purkayastha (2025). A new day gecko of the Cnemaspis podihuna (Reptilia: Gekkonidae) clade from northeastern India. Taprobanica 14(2): 107–120. https://doi.org/10.47605/tapro.v14i2.363
Schneider, J.G. (1797). Amphibiorum Physiologiae Specimen Alterum Historiam et Species Generis Stellionum seu Geckonum Sistens. Frankfurt (Oder): C.L.F. Aitzi, 54 pp.
Sengupta, D., C.G. Borah & J. Phukon (2019). Assessment of the reptilian fauna in the Brahmaputra Plains of two districts in Assam, India. Reptiles & Amphibians 26(1): 65–67. https://doi.org/10.17161/randa.v26i1.14346
Sengupta, S., J. Purakayasta, N. Anandan, M. Das, K.O. Chan & C. Murthy (2021). The odd one in: re-diagnosis and phylogenetic placement of the Assam Day Gecko, Cnemaspis assamensis Das & Sengupta, 2000 (Squamata: Gekkonidae). Zootaxa 5048(4): 581–593. https://doi.org/10.11646/zootaxa.5048.4.7
Sinha, B., K.P. Nath & S.D. Gurumayum (2021). Herpetofaunal diversity of Zoological Survey of India campus, Itanagar, Arunachal Pradesh, India. Records of the Zoological Survey of India 121(3): 411–418. https://doi.org/10.26515/rzsi/v121/i3/2021/156654
Smith, M.A. (1935). The Fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. Volume II. Sauria. Taylor and Francis, London, 185 pp.
Solanki, G.S. & A. Parida (2024). Study on re-establishment of herpetofauna in jhum fallow areas in Mizoram. In: Proceedings of the Zoological Society, Springer India, New Delhi: 1–9. https://doi.org/10.1007/s12595-024-00551-3
Tariang, A.D., F. Malsawmdawngliana, L. Biakzuala, H. Decemson, L. Muansanga, L. Rinsanga, M. Vabeiryureilai & H.T. Lalremsanga (2022). Confirmation on the occurrence of Calotes irawadi Zug, Brown, Schulte & Vindum, 2006 (Squamata: Agamidae) in Mizoram, northeastern India. Hamadryad 39(1 & 2): 88–94.
Thaosen, K., J. Purkayastha, M. Vabeiryureilai & H.T. Lalremsanga (2024). Sexual males in the parthenogenetic Hemidactylus cf. garnotii (Squamata: Gekkonidae) in Mizoram, India. Taprobanica 13: 124–131. https://doi.org/10.47605/tapro.v13i2.337
Uetz, P., P. Freed, R. Aguilar, F. Reyes & J. Hošek (eds.) (2025). The Reptile Database. http://www.reptile-database.org. Accessed on 20.vii.2025.
Wagner, P., F. Ihlow, T. Hartmann, M. Flecks, A. Schmitz & W. Böhme (2021). Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae). Bonn Zoological Bulletin 70(1): 141–171. https://doi.org/10.20363/BZB-2021.70.1.141
Wall, F. (1908). Remarks on Agamid lizard (Ptyctolaemus gularis). Journal of the Bombay Natural History Society 18: 505.
Web of Science (2025). https://www.webofscience.com. Accessed on 20.vii.2025.
Ulman, Y. & M. Singh (2021). Density, habitat associations and conservation status of Gekko gecko (Tokay gecko) in human-dominated landscapes around Nameri Tiger Reserve, Assam, India. Acta Ecologica Sinica 42(3): 188–197. https://doi.org/10.1016/j.chnaes.2021.03.004
Zhao, E. & S. Li (1984). A new species of Calotes (Lacertilia: Agamidae) from Xizang (Tibet) [in Chinese]. Acta Herpetologica Sinica 3(4): 77–78.
Zug, G.R., H.H.K. Brown, J.A. Schulte II & J.V. Vindum (2006). Systematics of the Garden Lizards, Calotes versicolor Group (Reptilia: Squamata: Agamidae), in Myanmar: central dry zone populations. Proceedings of the California Academy of Sciences 57(2): 35–68.
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