Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 January 2025 | 17(1): 26421–26425
ISSN 0974-7907 (Online)
| ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.9114.17.1.26421-26425
#9114 | Received 22
April 2024 | Final received 16 November 2024 | Finally accepted 12 January 2025
Survey
of Orthoptera in the Desert National Park, Rajasthan, India
Anshuman Pati 1, Indranil Paul
2 & Sutirtha
Dutta 3
1,2,3 Wildlife Institute of
India, Post Box #18, Chandrabani, Dehradun,
Uttarakhand 248001, India.
1,3 Academy of Scientific
and Innovative Research (AcSIR), Ghaziabad, Uttar
Pradesh 201002, India.
1 anshumanpati1996@gmail.com
(corresponding author), 2 29indranil@gmail.com, 3 sutirthadutta2016@gmail.com
Editor: Anonymity requested. Date of
publication: 26 January 2025 (online & print)
Citation: Pati, A., I. Paul & S. Dutta (2025). Survey of Orthoptera in the Desert
National Park, Rajasthan, India. Journal
of Threatened Taxa 17(1): 26421–26425. https://doi.org/10.11609/jott.9114.17.1.26421-26425
Copyright: © Pati et al. 2025. Creative Commons
Attribution 4.0 International License. JoTT allows
unrestricted use, reproduction, and distribution of this article in any medium
by providing adequate credit to the author(s) and the source of publication.
Funding: National Compensatory Afforestation Fund Management and Planning Authority (CAMPA), Government of India, and Theodore J. Cohn Research Fund for research award.
Competing interests: The authors declare no competing interests.
Acknowledgements: We are thankful to the director,
Wildlife Institute of India; dean (FWS); registrar and office staff of the Wildlife Institute of India, Dehradun for administrative support. The study was part of the Bustard Recovery Programme of the Wildlife Institute of India, funded by the National Compensatory Afforestation Fund Management and Planning Authority (CAMPA), Government of India. AP is thankful to the Theodore J. Cohn Research Fund for research award. Special thanks to the Bustard Recovery Project team for their guidance, encouragement and support during fieldwork. The authors are thankful to Mr. Devendar Genwa and Mr. Prakash Parihar for their assistance during the fieldwork. Special thanks to the Rajasthan Forest Department for providing the necessary work permission in the Desert National
Park. A special thanks to Dr. R. Swaminathan, Retired Professor of Entomology & Emeritus Scientist (ICAR) and Mrs. Tatiana Swaminathan, Honorary Museum Curator, Department of Entomology, Rajasthan College of Agriculture, Maharana Pratap University of Agriculture & Technology, Udaipur for their guidance in the specimen identification.
Abstract: Members of the insect
order Orthoptera comprising grasshoppers, locusts, and crickets form a dominant
and vital group of invertebrates in the arid environment. Orthopterans play an
important role in grassland ecosystems and their species diversity and abundance
in grasslands are much higher compared to those in the agricultural and
scrubland areas. We attempt to create a comprehensive list of Orthopteran
species from the Desert National Park (DNP) and its surrounding areas from the
Thar Desert landscape of Rajasthan. This area constitutes one of the largest
and few protected areas of arid biodiversity in India. In different enclosures
of DNP sweep net sampling was done to sample the focal species in different
seasons. 24 orthoptera species belonging to 20 genera and 11 subfamilies under
5 families were recorded.
Keywords: Grasshopper,
grassland, habitat, insect, season, sweep net, Thar Desert.
Deserts are found on
almost all the continents of the world and are characterized by dunes and
interdunal valleys. These regions experience extremely hot summers, cold
winters, and low, erratic rainfall. In India, the central parts of the desert
are occupied by grasslands. Grassland ecosystems are severely threatened by
agriculture and industrialization (White et al. 2000). Despite providing an
assortment of ecosystem services, supporting human livelihoods, and harbouring endemic wildlife, grasslands in India have been
largely ignored in development and conservation discourses (Dutta et al. 2010).
In grasslands, insect diversity is usually linked to plant species composition
and habitat structure (Roffey & Popov 1968).
Orthopteran insects,
especially grasshoppers, have a substantial importance in the ecology of
grassland ecosystems, being important primary herbivores and significantly
contributing to the diet of the Great Indian Bustard, an endangered bird of
Thar Desert (Dutta & Jhala 2021). Grasshoppers regularly
indicate habitat quality and change, so they are commonly regarded as potential
ecological indicator species for grasslands (Bazelet
& Samways 2011).
The order Orthoptera
is one of the significant insect orders, with about 29,530 species recorded
worldwide (Cigliano et al. 2024). There are about
1,274 species or subspecies belonging to 442 genera and 23 families recorded
from India (Chand et al. 2024). Some species of grasshoppers and locusts cause
considerable loss to vegetation in agricultural ecosystems in particular, in
many parts of the world due to their massive seasonal outbreaks (Klein et al.
2021).
In Thar Desert, the
Desert National Park (DNP) is an important grassland ecosystem with vast
expanses of grassland intermixed with shrubs and small trees. Earlier Rathore
(2004) reported only 10 species of Orthoptera from DNP. There is paucity of
information about the orthopteran species in the Thar landscape, specially the
DNP, being the only prominent protected area representative of India’s
north-western arid biogeographic zone. Therefore, the aim of our study was to
understand how many different type of orthopteran
species are there.
Methods
The Thar Desert
occupies nearly 385,000 km2 and about 9% of the area of India (Islam
& Rahmani 2004). Thar is occupied mainly by dry
open grassland or grassland interspersed with trees and shrubs with broad
topographic features like gravels, plains, sand-soil mix, dunes, and rocky
hillocks (Sharma & Mehra2009). The Desert National Park (DNP); actually a Wildlife Sanctuary encompasses about 3,162 km2
in the Jaisalmer District and another 1,262 km2 in the Barmer Districts of Rajasthan. Several areas within the
national park are protected by fencing where the human activity is restricted
to conserve the habitat for important wildlife species like the Great Indian
Bustard, where the vegetation is natural. Within the DNP several areas outside
the enclosures experience unrestricted human interference and movements, where
the vegetation of the few areas is natural and few areas are cultivated. The
study was carried out in different enclosures and unprotected agro-pastoral areas of DNP (Image 1). The size of the
enclosures are different, wherein, the minimum size of
the survey enclosures are 3 km2 and the maximum is 12 km2.
We surveyed these enclosures and adjoining unprotected areas in summer, monsoon
and winter seasons from June 2021 to December 2023. We visited the sampling
area multiple times in three monsoon, two winter and two summer seasons.
Sampling was done at
random points generated inside the study area. Sweep netting (Rudd & Jensen
1977) was done in a 50 x 4 m2 belt transect, following a standard
approach of about 100 strokes and the samples were collected in a container
after every 20 strokes. After that four adult individuals of different sexes of
each species were collected from the sample container. Preservation of the
collected specimen was done using wet and dry methods. For the dry preservation
insects were pinned and kept in a storage box and (Image 2) for the wet
preservation 70% ethanol were used. All the preserved insects were studied
under Nikon stereozoom microscope and identified
using taxonomic keys by Uvarov (1977). In addition,
we also used the species identification information from the Orthoptera Species
File (http://Orthoptera.SpeciesFile.org) last accessed on April 2024. The
collected specimens are deposited in the Great Indian Bustard Conservation
Breeding Center (Wildlife Institute of India), Pokhran, Rajasthan, India.
Results and Discussion
A total of 24 species
of Orthoptera were recorded from different enclosures (Protected) and
unprotected agro-pastoral areas. These species belong
to five families, 11 subfamilies, represent 20 genera, and four subgenera (Table
1). Notably 12 species were recorded in summer (April & May) while 24
species were observed during monsoon (July & August) and eight species in
winter (November & December). These species belong to four major types of
habitats, viz., grassland, scrubland, agricultural land, and barren land (Image
3).
The DNP is mainly
covered by grassland, and the dominant grass species include Dactyloctenium aegyptium,
Dactyloctenium scindicum,
Aristida depressa,
Cenchrus biflorus,
Lasiurus sindicus; apart from these, a few
shrub species are present comprising Aerva pseudotomentosa, Crotalaria burhia,
Dipterygium glaucum,
and Fagonia cretica
(Charan & Sharma 2016). During the monsoon all
the 24 species of Orthoptera were found which can be attributed to the
availability of green vegetation compared to other seasons when the vegetation
becomes predominantly dry. Acrididae was the most
abundant family recorded during the study followed by Pyrgomorphidae,
Tettigoniidae, Schizodactylidae,
and Tetrigidae. The present study adds to the
existing knowledge of the orthopteran fauna of DNP.
Table 1.
Orthoptera from Desert National Park during 2021–2023.
|
Family |
Subfamily |
Genera |
Subgenus |
Species |
Habitat |
|
Acrididae |
Acridinae |
Truxalis |
|
Truxalis indica (Bolívar, 1902) |
Grassland,
Scrubland |
|
Calliptaminae |
Acorypha |
|
Acorypha glaucopsis (Walker, 1870) |
Grassland |
|
|
Cyrtacanthacridinae |
Schistocerca |
|
Schistocerca gregaria (Forskål,
1775) |
Agriculture land,
Grassland |
|
|
Anacridium |
|
Anacridium rubrispinum Bey-Bienko,
1948 |
Grassland,
Scrubland |
||
|
Eyprepocnemidinae |
Heteracris |
|
Heteracris littoralis (Rambur, 1838) |
Scrubland |
|
|
Gomphocerinae |
Crucinotacris |
|
Crucinotacris decisa (Walker, 1871) |
Grassland |
|
|
Leva |
|
Leva indica (Bolívar, 1902) |
Grassland |
||
|
Ochrilidia |
|
Ochrilidia geniculata (Bolívar, 1913) |
Grassland |
||
|
|
Ochrilidia gracilis (Krauss, 1902) |
Grassland |
|||
|
|
Ochrilidia hebetata (Uvarov,
1926) |
Grassland |
|||
|
Oedopodinae |
Acrotylus |
|
Acrotylus humbertianus Saussure, 1884 |
Grassland |
|
|
|
Acrotylus longipes (Charpentier, 1845) |
Scrubland,
Grassland, Agriculture land |
|||
|
Oedaleus |
|
Oedaleus senegalensis (Krauss, 1877) |
Scrubland,
Grassland |
||
|
Scintharista |
|
Scintharista notabilis (Walker, 1870) |
Barren land |
||
|
Sphingonotus |
Neosphingonotus |
Sphingonotus (Neosphingonotus) paradoxus Bey-Bienko, 1948 |
Barren land |
||
|
Sphingonotus |
Sphingonotus (Sphingonotus) rubescens
(Walker, 1870) |
Barren land |
|||
|
Catantopinae |
Diabolocatantops |
|
Diabolocatantops pinguis (Stål,
1861) |
Scrubland |
|
|
Pyrgomorphidae |
Pyrgomorphinae |
Chrotogonus |
Chrotogonus |
Chrotogonus (Chrotogonus) trachypterus
(Blanchard, 1836) |
Barren land |
|
Poekilocerus |
|
Poekilocerus pictus (Fabricius, 1775) |
Scrubland |
||
|
Pyrgomorpha |
Pyrgomorpha |
Pyrgomorpha (Pyrgomorpha) bispinosa
Walker, 1870 |
Agriculture land |
||
|
Pyrgomorpha (Pyrgomorpha) conica (Olivier,
1791) |
Agriculture land |
||||
|
Tenuitarsus |
|
Tenuitarsus orientalis Kevan, 1959 |
Barren land |
||
|
Tetrigidae |
Tetriginae |
Paratettix Bolívar, 1887 |
|
Paratettix sp. |
Grassland |
|
Schizodactylidae |
Schizodactylinae |
Schizodactylus |
|
Schizodactylus monstrosus (Drury, 1773) |
Barren land |
|
Tettigoniidae |
Conocephalinae |
Euconocephalus |
|
Euconocephalus incertus (Walker, 1869) |
Grassland |
For
images - - click here for full PDF
References
Bazelet, C.S. & M.J.
Samways (2011). Identifying grasshopper bioindicators for habitat
quality assessment of ecological networks. Ecological Indicators 11(5):
1259–1269. https://doi.org/10.1016/j.ecolind.2011.01.005
Chand, D.S., S.K. Das, R. Chakraborty & H. Kumar
(2024). Checklist of Fauna of India: Arthropoda: Insecta:
Orthoptera. Version 1.0. Zoological Survey of India, 29 pp. https://doi.org/10.26515/Fauna/1/2023/Arthropoda:Insecta:Orthoptera
Charan, P.D. & K.C.
Sharma (2016). Floral diversity of Thar Desert of western Rajasthan,
India. Journal of Phytological Research 29(1 & 2): 55–71.
Cigliano, M.M., H. Braun,
D.C. Eades & D. Otte
(2024). Orthoptera Species File (version Apr 2024). In: Bánki,
O., Y. Roskov, M. Döring,
G. Ower, D.R. Hernández Robles, C.A. Plata Corredor, T. Stjernegaard
Jeppesen, A. Örn, L. Vandepitte,
D. Hobern, P. Schalk, R.E. DeWalt, K. Ma, J. Miller,
T. Orrell, R. Aalbu, J.
Abbott, R. Adlard & C. Aedo.
Catalogue of Life Checklist (Version 2024-04-26). Catalogue of Life.
Amsterdam, Netherlands. https://doi.org/10.48580/dg4lg-388
Dutta, S. & Y.V. Jhala
(2021). Devil is in the detail: behaviorally explicit habitat selection by the
Critically Endangered great Indian bustard. Endangered Species Research
45: 55–69. https://doi.org/10.3354/esr01126
Dutta, S., A.R. Rahmani
& Y.V. Jhala (2010). Running out of time?
The Great Indian Bustard Ardeotis nigriceps—status, viability, and conservation
strategies. European Journal of Wildlife Research 57(3): 615–625. https://doi.org/10.1007/s10344-010-0472-z
Islam, M.Z. & A.R. Rahmani
(2004). Important Bird Areas in India: Priority Sites for Conservation.
Indian Bird Conservation Network: Bombay Natural History Society and BirdLife International (UK), xviii+1133 pp.
Klein, I., N. Oppelt &
C. Kuenzer (2021). Application of
remote sensing data for locust research and management: a review. Insects
12(3): 233. https://doi.org/10.3390/insects12030233
Rathore, N.S. (2004). Conservation Area
Series. Zoological Survey of India, Kolkata, 19: 1–135.
Roffey, J. & G. Popov (1968). Environmental and behavioural processes in a desert locust outbreak. Nature
219(5153): 446–450.
Rudd, W.G. & R.L. Jensen (1977). Sweep Net and Ground
Cloth Sampling for Insects in Soybeans. Journal of Economic Entomology
70(3): 301–304. https://doi.org/10.1093/jee/70.3.301
Sharma, K.K. & S.P. Mehra
(2009). The Thar of Rajasthan (India): Ecology and Conservation of a Desert
Ecosystem, pp. 1–11. In: Sivaperuman, C., Q.H. Baqri, G. Ramaswamy & M. Naseema
(eds.). Faunal Ecology and Conservation of the Great Indian Desert.
Springer, Berlin, Heidelberg, xviii + 222 pp. https://doi.org/10.1007/978-3-540-87409-6_1
Uvarov, B. (1977). Grasshoppers and
Locusts Vol. II. Centre for Overseas Pest Research, London, 613 pp.
White, R., S. Murray
& M. Rohweder (2000). Pilot Analysis of
Global Ecosystems: Grassland Ecosystems. World Resources
Institute, Washington, D.C., 81 pp.