Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 January 2025 | 17(1): 26409–26416
ISSN 0974-7907 (Online)
| ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.9038.17.1.26409-26416
#9038 | Received 20
March 2024 | Final received 19 December 2024 | Finally accepted 26 December
2024
A
preliminary checklist of butterflies (Lepidoptera: Rhopalocera)
of Dhorpatan Valley, Dhorpatan
Hunting Reserve, Nepal
Kiran Rayamajhi 1, Bhaiya Khanal 2 & Prakash Chandra Aryal 3
1,3 GoldenGate International College (TU), Wisdom Tower, Trikuti Colony, Battisputali,
44600, Kathmandu, Nepal.
2 Nepal Bioheritage Forum for Resources Conservation (NBFRC), 25, Laligurans Galli, Gyaneshwor-1, 44600, Kathmandu, Nepal.
3 Environment
Protection and Study Center, Baneshwor, Prachin Marg, 44600, Kathmandu, Nepal.
1 rayamajhi82@gmail.com
(corresponding author), 2 baya2000@live.com, 3 pc.aryal@goldengate.edu.np
Editor: Anonymity requested. Date of
publication: 26 January 2025 (online & print)
Citation: Rayamajhi, K., B. Khanal
& P.C. Aryal (2025). A preliminary checklist of butterflies
(Lepidoptera: Rhopalocera) of Dhorpatan
Valley, Dhorpatan Hunting Reserve, Nepal. Journal of Threatened Taxa 17(1): 26409–26416. https://doi.org/10.11609/jott.9038.17.1.26409-26416
Copyright: © Rayamajhi et al. 2025. Creative Commons
Attribution 4.0 International License. JoTT allows
unrestricted use, reproduction, and distribution of this article in any medium
by providing adequate credit to the author(s) and the source of publication.
Funding: The Study and Research Aid Program for the fiscal year 2077/078 BS (2020/21 AD) by Department of Environment (DoE), Ministry of Forests and Environment, Nepal (MoFE).
Competing interests: The authors declare no competing interests.
Acknowledgements: The authors express their sincere gratitude to the Department of Environment (DoE), Ministry of Forests and Environment, Nepal (MoFE), for providing grant support for this research under the Study and Research Aid Program for the fiscal year 2077/078 BS (2020/21 AD). This study was conducted as a part of the first author’s master’s thesis. We are also grateful to Mr. Purushottam Sharma, former Senior Conservation Officer of Dhorpatan Hunting Reserve (DHR), for his encouragement and support. Furthermore, we extend our heartfelt thanks to the dedicated officers of the reserve for their valuable assistance and cooperation during the fieldwork period
Abstract: Butterfly species
were studied in the Dhorpatan Valley, situated in the
western region of Nepal, during the pre-monsoon and monsoon seasons of 2021 and
2022. This preliminary study documented forty-three species of butterflies from
five families Lycaenidae, Pieridae,
Nymphalidae, Papilionidae,
and Hesperiidae. A purposive sampling method was
applied to locate various butterfly species across elevations ranging
2,846–4,000 m. This study recorded Polyommatus
nepalensis, an endemic species and Polyommatus stoliczkanus,
a rare species. No reports have been made of this endemic butterfly from
eastern region of Nepal.
Keywords: Distribution,
elevation, endemic, family, habitat, plots, rare, sampling, seasons, species.
The IUCN Butterfly
and Moth Specialist Group estimates that there are approximately 18,000
butterfly species and 160,000 moth species globally. Insects make up more than
half of the world’s faunal diversity, which plays a crucial role in the
functioning of the earth’s ecosystem. Butterflies are the best-studied group
throughout Nepal (Smith 1994, 1997). These insects are widely distributed in
Nepal and 18% of butterflies in the mid-hills are threatened (BPN 1996; Bhusal & Khanal 2008; Thapa
& Bhusal 2009). The country’s various
bio-climatic zones reflect the niches for a wide variety of flowers that offer
ideal habitats for butterfly diversity.
Six-hundred-and-sixty
species under 263 genera of butterflies have been reported in Nepal (Smith
2010). Based on Smith (2010) and Subedi et al.
(2021), the most recent count of butterfly species in Nepal is 678. Since, no
previous studies on invertebrate fauna, such as butterflies and moths, have
been conducted in this region, this research aims to provide important
information on the butterfly species inhabiting the higher elevations of Nepal.
This preliminary checklist of butterfly species in this region could also help
explore habitat preferences, particularly in relation to larval host plants
such as Rumex nepalensis,
Berberis aristata,
Duchesnea indica,
Anaphalis spp., and Pedicularis
spp. This information contributes to initiating conservation efforts,
particularly in protected areas of Nepal like Dhorpatan
Hunting Reserve.
Study area
Dhorpatan Valley is the part of Dhorpatan
Hunting Reserve (DHR) located in Baglung District,
Gandaki Province, western region of Nepal (28.490 0N, 83.027 0E)
(Figure 1, Image 1). This reserve was established to protect a variety of
Himalayan flora & fauna, represented by alpine, sub-alpine, and high
temperate vegetation. According to the DHR (2019), the features include
extensive highland pastures mostly above 3,800 m and east-west ridges that make
north and south slopes suitable for summer and winter habitats. Forest covers
different vegetations like blue pine Pinus wallichiana,
fir Abies spectabilis,
rhododendron (Rhododendron ferrugineum,
R. campanulatum), Hemlock Tsuga
dumosa, Birch Betula utilis,
Juniper Juniperus indica,
Spruce Picea smithiana,
and oak Quercus semicarpifolia. This is the
only one hunting reserve of Nepal that supports sport hunting of Jharal Hemitragus jemlahicus and Blue Sheep Pseudois
nayaur. The map of the study area was drawn using
Arc GIS 10.5.1. The survey of butterfly was conducted in the Dhorpatan Valley of the Dhorpatan
Hunting Reserve (DHR), where the altitude ranges 2,846–4,000 m. The diverse
physiographic conditions of this valley, including seasonal meadows, various
water bodies, and a wide range of shrubs & herbs, provide preferred
habitats and food for butterflies & moths, supporting the pollination of
floral species, and contributing to overall ecosystem health &
biodiversity.
Materials and Methods
Butterfly species
were recorded from 50 plots with a plot size of 50 × 20 m2 in the
study area of Dhorpatan Valley. This research was
conducted over two months during the pre-monsoon season (25 March–23 April, 2021) and the monsoon season (19 June–18 July, 2022).
The Pollard (1977) method was followed, where butterflies were surveyed between
1045 h and 1545 h on sunny days with temperatures (>17 °C, or 13–17 °C). The
butterflies were observed and recorded from the sampling plots for five hours
per day throughout the study period using purposive sampling. Most of the
species were photographed using a Nikon D-7500 camera, and only a few confusing
or unidentifiable species were collected. Different relevant literatures (Khanal & Smith 1997; Smith 2011) were consulted for
identification.
Results and Discussion
This study records 43
species of butterflies (Images 2–47), categorization based on their
availability relative to the total number of individuals recorded in this
region. All the recorded species belong to 32 genera and five families, which
include Lycaenidae (15 species), Pieridae
(8 species), Nymphalidae (14 species), Papilionidae (4 species), and Hesperiidae
(2 species). Single individuals of Parnassius
hardwickii, Papilio
protenor, Everes
lacturnus, Parantica
sita, Argynnis
kamala, and Neptis hylas
were recorded above 3,000 m in the Dhorpatan Valley.
Inomata (1998) published an account of Parnassius
of Nepal which includes 12 species from the Himalayan region. Only few studies
provide information on the population status of Apollo species and their
habitat preferences in different geographical regions of the world (Ali et al.
2019). Aglais cashmirensis
was noted with the highest abundance records during the whole period
of survey. This is one of the most common species, found in a wide range of
habitats in India (Haribal 1990). Polyommatus nepalensis, an
endemic species, and Polyommatus stoliczkanus, a rare species, were generally recorded
as common in this geographic region.
We observed that the
monsoon season, typically from June to July, is an optimal time for monitoring
butterfly diversity in high-altitude regions like Dhorpatan
Valley. The pre-monsoon season is typically dry so, few individuals of species
like Polyommatus icarus,
Polyommatus nepalensis,
Polyommatus stoliczkanus,
Vanessa indica, Junonia
iphita, Parnassius
hardwickii, Everes argiades, and Udara albocaerulea
were mostly seen. Within this protected area, most butterfly species were
recorded at lower elevations. The diverse habitats of this reserve support a
wide variety of flora, fauna, and invertebrates, including butterflies, moths,
and bees. As pollinators, butterflies & bees play a crucial role in
maintaining floral diversity and enhancing the overall health of the ecosystem
(Potts et al. 2010). This preliminary study fills a significant knowledge gap
by providing the first comprehensive assessment of butterfly species in this
region. The baseline data generated by this research will be instrumental in
guiding future studies and conservation initiatives.
Table 1. A
preliminary checklist of butterfly species found in the Dhorpatan
Valley of Dhorpatan Hunting Reserve, Nepal. The list
includes family, names of the species, and their local status.
|
Family |
Scientific name |
Common name |
Local status |
|
1. Pieridae |
1. Catopsilia pyranthe
Linnaeus, 1758 |
Mottled Emigrant |
C |
|
2. Colias erate Esper, 1805 |
Eastern Pale
Clouded Yellow |
C |
|
|
3. Colias fieldii Ménétriés, 1855 |
Dark Clouded Yellow |
C |
|
|
4. Delias belladonna Fabricius,
1793 |
Hill Jezebel |
C |
|
|
5. Eurema hecabe Linnaeus,
1758 |
Oriental Common
Grass Yellow |
C |
|
|
6. Gonepteryx nepalensis Linnaeus, 1758 |
Common Brimstone |
C |
|
|
7. Pieris brassicae Linnaeus, 1758 |
Large Cabbage White |
C |
|
|
8. Pieris canidia Sparrman, 1768 |
Indian Cabbage
White |
C |
|
|
2. Nymphalidae |
9. Aglais cashmirensis Kollar, 1844 |
Indian
Tortoiseshell |
FC |
|
10. Argynnis kamala Moore, 1857 |
Common Silverstripe |
FC |
|
|
11. Aulocera brahminus Blanchard,
1853 |
Narrow-banded Satyr |
UC |
|
|
12. Lethe sidonis Hewitson, 1863 |
Common Woodbrown |
C |
|
|
13. Issoria issaea Doherty, 1886 |
Queen of Spain
Fritillary |
C |
|
|
14. Junonia orithya Linnaeus,
1758 |
Blue Pansy |
C |
|
|
15. Lasiommata schakra Kollar, 1844 |
Common Wall |
C |
|
|
16. Melitaea arcesia Bremer,
1861 |
Blackvein Fritillary |
C |
|
|
17. Neptis hylas Linnaeus, 1758 |
Common Sailor |
VC |
|
|
18. Parantica sita Kollar, 1844 |
Chestnut Tiger |
LC |
|
|
19. Junonia iphita Cramer, 1779 |
Chocolate Pansy |
C |
|
|
20. Rhaphicera moorei Butler,
1867 |
Small Tawny Wall |
C |
|
|
21. Vanessa cardui Linnaeus, 1758 |
Painted Lady |
C |
|
|
22. Vanessa indica Herbst, 1794 |
Indian Red Admiral |
VC |
|
|
3. Lycaenidae |
23. Celastrina argiolus
Linnaeus, 1758 |
Hill Hedge Blue |
C |
|
24. Celastrina gigas Hemming,
1928 |
Silvery Hedge Blue |
C |
|
|
25. Everes lacturnus Godart, 1824 |
Indian Cupid |
FC |
|
|
26. Everes argiades Pallas,
1771 |
Tailed Cupid |
C |
|
|
27. Heliophorus epicles Godart, 1824 |
Purple Sapphire |
C |
|
|
28. Heliophorus sena Kollar, 1844 |
Sorrel Sapphire |
UC |
|
|
29. Lampides boeticus Linnaeus,
1767 |
Pea Blue |
C |
|
|
30. Leptotes plinius Fabricius, 1793 |
Zebra Blue |
UC |
|
|
31. Lycaena panava Westwood,
1852 |
White-bordered
Copper |
LC |
|
|
32. Lycaena phlaeas Linnaeus,
1761 |
Common Copper |
C |
|
|
33. Polyommatus icarus
Rottemburg, 1775 |
Common Blue |
UC |
|
|
34. Polyommatus nepalensis
Forster, 1961 |
Nepal Meadow Blue |
E |
|
|
35. Polyommatus stoliczkanus
C. & R.Felder, 1865 |
Himalayan Meadow
Blue |
R |
|
|
36. Pseudozizeeria
maha Kollar, 1844 |
Pale Grass Blue |
FC |
|
|
37. Udara albocaerulea Moore, 1879 |
Himalayan Albocerulean |
C |
|
|
4. Papilionidae |
38. Graphium cloanthus
Westwood, 1841 |
Glassy Bluebottle |
UC |
|
39. Papilio machaon Linnaeus,
1758 |
Common Yellow
Swallowtail |
C |
|
|
40. Papilio protenor
Cramer, 1775 |
Spangle |
C |
|
|
41. Parnassius hardwickii Gray,
1831 |
Common Blue Apollo |
UC |
|
|
5. Hesperiidae |
42. Pelopidas mathias Fabricius, 1798 |
Small Branded Swift |
C |
|
43. Caltoris cahira austeni (Moore,
1883) |
Colon swift |
C |
C—Common |
UC—Uncommon | LC—Locally Common | FC—Fairly Common | R—Rare | E—Endemic.
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figure & images - - click here for full PDF
References
Ali, M., V. Kumar & K. Chandra (2019). Butterfly Diversity
in different Trans-Himalayan Habitats of Ladakh,
Jammu & Kashmir, India. Proceedings of the 2nd Himalayan
Researchers Consortium – Vol. 3. Dehradun, Uttarakhand, 22 pp.
Bhusal, D.R. & B. Khanal (2008). Seasonal and Altitudinal Diversity of
Butterflies in Eastern Siwalik of Nepal. Journal of Natural History Museum
23: 82–87. https://doi.org/10.3126/jnhm.v23i0.1843
BPN (1996). Biodiversity Profile
of Nepal with special reference to protected area. Dept. of National Park and
Wildlife Conservation HMG/Nepal, 51–60 pp.
DHR (2019). Dhorpatan
Hunting Reserve Management Plan (2076/77-2080/81) Dhorpatan
Hunting Reserve Office, Dhorpatan, Baglung, Nepal.
Haribal, M. (1990). The behavior of the
Indian Tortoiseshell Butterfly Aglais (Vanessa)
cashmiriensis (Kollar)
in the Himalayas. Journal of the Bombay Natural History Society 87(1):
163–164.
Inomata, T. (1998). Parnassinae
from Nepal. Moths of Nepal, pp. 311–314. In: Haruta,
T. (ed.), Part 5. The Japan Hetrocerist’s Society,
Tokyo.
Khanal, B. & C. Smith
(1997). Butterflies of Kathmandu Valley. Craftsman Press, Bangkok, 115
pp.
Pollard, E. (1977). A method for
assessing changes in abundance of butterflies. Biological Conservation
12: 115–134. https://doi.org/10.1016/0006-3207(77)90065-9
Potts, S.G., J.C. Biesmeijer,
C. Kremen, P. Neumann, O. Schweiger & W.E. Kunin (2010). Global pollinator declines: trends, impacts and
drivers. Trends in Ecology & Evolution 25(6): 345–353. https://doi.org/10.1016/j.tree.2010.01.007
Smith, C. (1994). Butterflies of
Nepal (Central Himalaya). Revised Edition. T.C. Majupariya
and Craftman Press, Bangkok, 352 pp.
Smith, C. (1997). Butterflies of
Royal Chitwan National Park, Nepal. Tecpress
Books, Bangkok, 80 pp.
Smith, C. (2010). Butterflies of
Nepal. Himalayan Nature, 184 pp.
Smith, C. (2011). An Illustrated
Checklist of Nepal’s butterfly. Craftsman Press, Bangkok, 131 pp.
Subedi, N., S. Bhattarai,
M.R. Pandey, R. Kadariya, S.K. Thapa, A. Gurung, A. Prasai, S. Lamichhane, R. Regmi, M. Dhungana, P.R. Regmi, R.P. Paudel, B. Kumpakha, B. Shrestha, B. Gautam, R. Baral,
U. Poudel, S. Yadav, S. Pariyar & B.R. Lamichhane (2021). Report on Faunal
Diversity in Chure Region of Nepal. President Chure-Terai Madhesh Conservation
Development Board and National Trust for Nature Conservation. Kathmandu, Nepal.
Thapa, G. & D.R. Bhusal (2009). Species Diversity and Seasonal Variation of
Butterfly Fauna in Thankot and Syuchatar
VDC of Kathmandu Valley, Nepal. Journal of Natural History Museum 24(1):
9–15. https://doi.org/10.3126/jnhm.v24i1.2229