Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2025 | 17(5): 26985–26998

 

ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print) 

https://doi.org/10.11609/jott.9532.17.5.26985-26998

#9532 | Received 03 December 2024 | Final received 26 February 2025 | Finally accepted 16 May 2025

 

 

Plant composition and species diversity in Delhi NCR of India

 

Amit K. Tripathi 1  & Jyoti K. Sharma 2        

 

1,2 School of Natural Sciences, Shiv Nadar University, Greater Noida, Uttar Pradesh 201314, India

1 amit.tripathi@snu.edu.in (corresponding author) 2 jyotikumarsharma@gmail.com

 

                                                        

Editor: A.J. Solomon Raju, Andhra University, Visakhapatnam, India.          Date of publication: 26 May 2025 (online & print)

 

Citation: Tripathi, A.K. & J.K. Sharma (2025). Plant composition and species diversity in Delhi NCR of India. Journal of Threatened Taxa 17(5): 26985–26998. https://doi.org/10.11609/jott.9532.17.5.26985-26998

  

Copyright: © Tripathi & Sharma 2025. Creative Commons Attribution 4.0 International License. JoTT allows unrestricted use, reproduction, and distribution of this article in any medium by providing adequate credit to the author(s) and the source of publication.

 

Funding: All the logistics and support received from Shiv Nadar University, Delhi NCR.

 

Competing interests: The authors declare no competing interests.

 

Author details: Dr. Amit K. Tripathi is currently working as a postdoctoral research associate in School of Natural Sciences, Shiv Nadar University, Delhi NCR. With over ten years of research experience, His main areas of interest are forest ecology, Angiosperm taxonomy, and biodiversity documentation and conservation. Dr. Jyoti K. Sharma, former director of Kerala Forest Research Institute (KFRI), has been a professor and HOD of Center for Environmental Sciences and Engineering, School of Natural Sciences, Shiv Nadar University (Delhi NCR), from 2013 to 2023. He has over 40 years’ of research experience in the field of forestry, natural resource management, biodiversity assessment & conservation.

 

Author contributions: AKT carried out the field survey, compiled data, captured photographs and prepared the manuscript. JKS, as a principal investigator, mentored the whole research work, reviewed, and edited the manuscript. Both authors approved the final submitted manuscript.

 

Acknowledgements: This work is part of the research project “Biodiversity documentation of Chithara Gram Panchayat, Dadri Block Panchayat, Tehsil Dadri, District Gautam Buddha Nagar, Uttar Pradesh and preparation of a Biodiversity Register” funded by Shiv Nadar University, Delhi NCR, India. The authors are grateful to the Vice-Chancellor of Shiv Nadar University for her unstinted support during the course of this investigation.

 

 

Abstract: The present study provides a first major step in documenting and conserving the local plant diversity in Delhi NCR, which witnessed high species richness within a relatively small, localized area and harbors many high-value medicinal plants used in Ayurveda and other Indian traditional systems of medicine. The local hotspot areas that serve as a unique ecosystem for flora are being depleted day by day as a result of anthropogenic pressure. Hence, there is a need to conserve the species hotspots involving local communities who always interact with these ecosystems. The present study documents 272 plant species belonging to 204 genera and 69 families from the Delhi NCR of India. The aquatic and terrestrial vegetation in the study area was surveyed frequently throughout the year in different seasons, and plant collections were made. The majority of species documented were flowering plants, which comprised 216 dicots, 53 monocots, and three species belonged to lycophytes and fern groups. Fabaceae was the most dominant family with 39 plant species, followed by Poaceae (31 species), Asteraceae (25 species), Convolvulaceae (14 species), and Amaranthaceae and Malvaceae (12 species each). Genus Ipomoea is recorded to have the highest number of species (7), followed by Euphorbia and Cyperus, having five species each. Overall, the herbaceous community of species was greater in numbers than other life forms.

 

Keywords: Biodiversity documentation, diverse habitats, high-value herbs, local flora, rare plants, sustainable utilization, village panchayat, western Uttar Pradesh, wild plants.

 

 

INTRODUCTION

 

Floristic surveys and subsequent documentation play an important role in the conservation of biodiversity (Sharma et al. 2018). Repeated surveys in any area in different seasons increase the chances of documenting rare and unique species, but the absence of knowledge about the species present and taxonomic identification of individual species makes it difficult to assess the richness of plant diversity.

Western Uttar Pradesh, within Delhi NCR (National Capital Region) has some demographic, economic, and cultural patterns that are distinct from other parts of Uttar Pradesh, and more closely resemble those of Haryana & Rajasthan states. A major part of western Uttar Pradesh falls within National Capital Region (NCR) of India and is experiencing rapid economic growth due to intensive agriculture and infrastructure development; hence, it is imperative to document and conserve the floristic wealth of these areas before the plant species are threatened and lost (Sharma et al. 2018).

Very few studies (Singh 1969; Vardhana 2007; Ahamed & Gupta 2010; Shishodia 2013; Malik 2015) are available supporting the floristic studies of western Uttar Pradesh, which come under Delhi NCR. The first major step in documenting and conserving the local plant diversity is the survey of the flora of that region. There is a need for proper inventory and documentation of all plants available in the western Uttar Pradesh NCR region, which will help formulate a much-needed strategy for conservation.

The present study has been carried out in Delhi NCR of western Uttar Pradesh. The study provides baseline information on floristic diversity with some unique, rare, and new species record that will help in updating the flora of India. The study also highlights many high-value medicinal herbs growing in the area. The present work is the first exhaustive study of its kind in Delhi NCR and is vital for the conservation of local flora and also for increasing awareness among local people.

 

 

MATERIAL AND METHODS

 

Study area

The study area is the Chithara Village Panchayat including the campus of Shiv Nadar University, Gautam Buddha Nagar District in western Uttar Pradesh (Figure 1); it is the eastern part of Delhi NCR. This region falls in the Gangetic plains, which are one of the most fertile regions of the country and equally rich in wild plant diversity. Geographically, Chithara lies between 28.516–28.55 °N and 77.55–77.58 °E, and is one of the 56 Panchayats under Dadri Block Panchayat/Tehsil and covers an area of 7.7 km2. The village has a population of 7,656 of which 4,098 are males while 3,558 are females as per Population Census 2011. The village is divided into mohallas largely based on communities. Gujjar and Jatav communities are prominent in the village.

Chithara is a sub-urban area with a semi-arid climate. The climate is typically monsoonal with three distinct seasons, namely, summer (March–June), rainy (July–September), and winter (October–February). It experiences the hottest weather in June, with maximum temperature around 450C. The coldest month is January, with minimum temperature around 50C. August is usually the wettest month during the rainy season.

The soil is fertile alluvial, whose pH ranges between 7.64–9.38 with reasonably good maximum water holding capacity. Nutrient-wise the soil contained organic carbon (0.299–0.400%), which is within the desirable range of soils of upper Gangetic Plain. The village is surrounded by agricultural fields and several wetlands. The wild date palm tree, Phoenix sylvestris (L.) Roxb., is scattered as individual trees, in small groups, and in large groves. The village has an agriculture-based economy, mainly having wheat, and paddy cultivation.

 

Methods

The biodiversity documentation of Chithara Village Panchayat was carried out during 2015–2019.  Field surveys covered different types of habitats such as agricultural fields, canal bunds, roadsides, village streets, wild date palm groves, wetlands, wastelands, grasslands (Sharma et al. 2018). The aquatic and terrestrial vegetation of the area was surveyed frequently throughout the year, covering different seasons. Photographs of each species were taken in the field for identification purposes. The plants were identified with the help of available floras (Duthie 1960; Maheshwari 1966; Sharma & Dhakre 1995; Vardhana 2007) and herbaria of the Botanical Survey of India, Northern Regional Office, Dehradun, and the Forest Research Institute, Dehradun. The specimens of identified plants were deposited in the BSI herbarium, Dehradun and CESE Department, Shiv Nadar University (SNU/CESE). The taxonomic identity of the plant specimens was verified from POWO (2024). The medicinal properties and medicinal uses of each plant species were studied, and among the list, 10 highly important medicinal plants were identified, which are known for their outstanding medicinal properties and are used in Ayurveda and other traditional systems of medicine (Sharma et al. 2018). Plant species with very restricted distribution were considered locally rare.

 

 

RESULTS AND DISCUSSION

 

The present study documented a total of 272 plant species belonging to 204 genera and 69 families. The majority of species were flowering plants (269 plants: dicots—216 species, monocots—53 species) and only three species belonged to lycophytes and fern categories (Table 1).

In terms of percentage, 79% of species were dicotyledons, 20% were monocotyledons, and 1% were lycophytes & fern categories (Figure 2). There was a preponderance of tall grasses such as Saccharum spontaneum L., Saccharum bengalense Retz., and Phragmites karka (Retz.) Trin. ex Steud., particularly in the wetland areas, which is very typical of uncultivated lands in the region of Gangetic plains.

Previously published study reveals that the flora of Delhi had been extensively explored by Maheshwari (1963) who reported a total of 531 species from the region (Table 2). The present study reported 272 plant species (Table 1), which look-like small in number with respect to earlier work, but in terms of the geographic area, it is a very high proportion, which is only 7.7 km2 in comparison to large geographical area of NCT of Delhi (1,483 km2).

Documentation of such a large number of species, which constitutes about 50% of flora of Delhi in such a small geographical area, clearly showed that the diverse habitats such as agricultural fields, roadsides, village streets, wetlands, canal bunds, wastelands, grasslands, and wild date palm groves, in the study area contributed to the rich plant diversity coupled with fertile soil conditions. The study area recorded many medicinal herbs growing as weeds (Tripathi et al. 2019). Sometimes, medicinal herbs growing as weeds contribute significantly to enhance the local plant diversity (Tripathi et al. 2020). The study area witnessed some prominent groves of wild date palms P. sylvestris associated with some other trees, herbs, and shrubs.

The floristic diversity in Delhi, and the National Capital Region (NCR) under western Uttar Pradesh was studied from time to time (Maheshwari 1963; Singh 1969; Vardhana 2007; Ahamed & Gupta 2010; Shishodia 2013; Malik 2015) (Table 2). Compared to all these studies carried out in a specific area/district , the present study has been carried out in a village panchayat in Delhi NCR encompassing high plant diversity.

 

Family and genera dominance

With respect to families, Fabaceae was the most dominant with 39 plant species followed by Poaceae (31 species), Asteraceae (25 species), Convolvulaceae (14 species), and Amaranthaceae & Malvaceae (12 species each) (Figure 3). Euphorbiaceae & Solanaceae contributed eight species each, while Cyperaceae & Brassicaceae had seven, and six species, respectively. These families contributed approximately 60% of total plants. The dominance of these families was also reported by various authors (Duthie 1960; Vardhana 2007). Out of 272 vascular plant species, dicotyledons were represented by 56 families, 160 genera, 216 species; monocotyledons by 10 families, 41 genera, 53 species, and lycophytes & fern by three families, three genera, and three species (Figure 4). Among the dicots, the most dominant family was Fabaceae represented by 24 genera, and 39 species. Of the 10 families represented in monocotyledons, Poaceae was the most versatile family with the highest number of species belonging to 25 genera and 31 species, contributing more than 50 % species of the total monocots (Table 1). Genus Ipomoea had the highest number of seven species. This was followed by Euphorbia (Euphorbiaceae) and Cyperus (Cyperaceae) having five species each. There were 46 genera under 23 families with at least two species each. Fourteen families had a single species each while six families had two species each.

 

Medicinally important plants

The study area witnessed many high-value medicinal plants that play a specific role in ayurveda and other systems of medicine. The list includes Bacopa monnieri (L.) Wettst. (Brahmi), Boerhavia diffusa L. (Punarnava), Cannabis sativa L. (Bhang), Cissampelos pareira L. (Patha), Coccinia grandis (L.) Voigt (Kundru), Eclipta prostrata (L.) L. (Bhringaraj), Euphorbia hirta L. (Bari Dudhi), Justicia adhatoda L. (Arus), Sida cordifolia L. (Bala), Solanum americanum Mill (Makoy), Tribulus terrestris L. (Gokhru), and Withania somnifera (L.) Dunal (Ashwagandha) which were found to be very important as they possess medicinal properties for treating a large number of diseases in traditional system of medicines in India and other parts of the world (Image 1). The whole plant, including root, stem, and leaves is utilized traditionally for curing various ailments such as abdominal pain, anxiety, cold, cough, bronchitis, asthma, diarrhea & dysentery, dull memory, epilepsy, eye troubles, fever, inflammation, jaundice, rheumatism, skin diseases, cardiovascular problems, sexual dysfunction, and various urinary disorders (Sharma et al. 2018). These plants have been reported to contain numerous important phytochemicals such as alkanes, amino acids, flavonoids, organic acids, polyphenolic compounds, steroids, tannins, and terpenes, which qualify their various medicinal properties (Tripathi et al. 2020). Certain plant species have been reported to yield potent drugs such as vasicine and vasicinone in Justicia adhatoda, punarnavine in Boerhavia diffusa, bacosoids in Bacopa monnieri, protodioscin in Tribulus terrestris and withanolides in Withania somnifera (Sharma et al. 2018). Sustainable utilization and conservation of these multi-purpose herbs for medicinal purposes is inevitable. Globally, traditional medicine, mostly herbal medicine, is known as a major healthcare provider in rural as well as remote areas, and a large number of people in developing, and underdeveloped countries depend on such medicine for their health (Sen & Chakraborty 2017).

 

Rare and newly recorded species

The species whose occurrence was very restricted and found only at one or two sites were considered rare species. Among the list, three species, such as Crateva adansonii subsp. odora (Buch.-Ham.) Jacobs, Mitragyna parviflora (Roxb.) Korth., and Streblus asper Lour. were rare occurrences in the study area.

The survey documented a few unique species, which were recorded for the first time in this region, such as Rumex hypogaeus T.M.Schust. & Reveal (earlier Emex australis Steinh.), Phyllanthus tenellus Roxb., Equisetum ramosissimum Desf., and Symphyotrichum squamatum (Spreng.) G.L.Nesom. R. hypogaeus belonging to family Polygonaceae, was found growing gregariously in a few places on the SNU campus of the Chithara village and it was a new record for Uttar Pradesh and upper Gangetic plains (Tripathi et al. 2018). P. tenellus (Phyllanthaceae), an annual herb that is common in southern India, has expanded its distribution to northern India. E. ramosissimum (ssp. ramosissimum and ssp. debile) was also recorded from the study area; both subspecies were found for the first time from western Uttar Pradesh and NCR Delhi (Sharma et al. 2018). In addition, S. squamatum, an erect herb of the Asteraceae, was also recorded for the first time in India from the study area (Tripathi & Sharma 2019).

As a result of repeated surveys in different seasons in heterogeneous habitats of the study area, including agricultural fields, wetlands, canal bunds, wild date palm groves, and roadsides, it became possible to document many rare, and new record species. Due to anthropogenic pressure, the local hotspot areas for biodiversity, such as wild date palm groves & wetlands, which provide habitat to numerous species, are being depleted day by day. Therefore, there is a need to conserve the species hotspots and to create general awareness among local communities.

 

 

CONCLUSION

 

As a first step towards conservation and sustainable utilization of biodiversity, work on biodiversity documentation was undertaken in Chithara Village Panchayat, where so far no efforts have been made to prepare the People’s Biodiversity Register as mandated in the Biological Diversity Act (2002). The extensive floristic survey of the village resulted in high species richness, which represents the diversity within a relatively small, localized area with potentially varied habitats within it. A rise in anthropogenic disturbances could threaten important plant habitats in the future, hence, local hotspot areas, such as wetlands, and wild date palm groves that serve as a unique ecosystem for flora and fauna, should be restored and community-based conservation and management of flora need to be emphasized since local communities always interact with village ecosystems. The study indicates that rural India remains rich in wild plant diversity, including high-value medicinal herbs, and may still harbor many unrecorded species. Hence, long-term, extensive documentation of flora at the village panchayat level should be prioritized.

 

Table 1. List of the plant species in Chithara Village Panchayat, Delhi NCR.

 

Botanical name

Family

Vernacular/ Common name

Herbarium catalogue no.

 

Herbs

 

 

 

1

Achyranthes aspera L.

Amaranthaceae

Chirchita, Latjira

SNU/CESE/188

2

Acmella paniculata (Wall. ex DC.) R.K.Jansen

Asteraceae

Toothache Plant

SNU/CESE/277

3

Ageratum conyzoides (L.) L.

Asteraceae

Billygoat-weed

SNU/CESE/073

4

Ageratum houstonianum Mill.

Asteraceae

Floss Flower

SNU/CESE/422

5

Alhagi maurorum Medik.

Fabaceae

Camelthorn-bush

SNU/CESE/435

6

Alternanthera paronychioides A.St.-Hil.

Amaranthaceae

Smooth Chaff Flower

SNU/CESE/392

7

Alternanthera philoxeroides (Mart.) Griseb.

Amaranthaceae

Alligator Weed

SNU/CESE/285

8

Alternanthera pungens (Kunth)

Amaranthaceae

Chaff-Flower

SNU/CESE/101

9

Alternanthera sessilis (L.) R.Br. ex DC.

Amaranthaceae

Sessile Joyweed

SNU/CESE/152

10

Alysicarpus monilifer (L.) DC.

Fabaceae

Necklace-pod Alyce Clover

SNU/CESE/130

11

Alysicarpus vaginalis (L.) DC.

Fabaceae

Alyce Clover

SNU/CESE/159

12

Amaranthus viridis L.

Amaranthaceae

Jangali Chaulai

SNU/CESE/002

13

Ammannia baccifera L.

Lythraceae

Blistering Ammannia

SNU/CESE/236

14

Anisomeles indica (L.) Kuntze

Lamiaceae

Kala Bhangra

SNU/CESE/251

15

Arabidopsis thaliana (L.) Heynh.

Brassicaceae

Thale Cress

SNU/CESE/316

16

Arenaria serpyllifolia L.

Caryophyllaceae

Thyme-leaved Sandwort

SNU/CESE/362

17

Argemone mexicana L.

Papaveraceae

Satyanashi

SNU/CESE/038

18

Artemisia scoparia Waldst. & Kitam.

Asteraceae

Redstem Wormwood

SNU/CESE/168

19

Avena sterilis L.

Poaceae

Jangali Jai

SNU/CESE/325

20

Bacopa monnieri (L.) Wettst.

Plantaginaceae

Water Hyssop (Brahmi)

SNU/CESE/220

21

Barleria prionitis L.

Acanthaceae

Vajradanti

SNU/CESE/300

22

Bidens pilosa L.

Asteraceae

Black-jack

SNU/CESE/071

23

Blainvillea acmella (L.) Philipson

Asteraceae

Para Cress Flower

SNU/CESE/208

24

Blumea laciniata (Wall. ex Roxb.) DC.

Asteraceae

Cutleaf Blumea

SNU/CESE/423

25

Blumea obliqua (L.) Druce

Asteraceae

Common Floss Flower

SNU/CESE/269

26

Blumea viscosa (Mill.) V.M.Badillo

Asteraceae

Sticky Blumea

SNU/CESE/427

27

Boerhavia diffusa L.

Nyctaginaceae

Punarnava

SNU/CESE/341

28

Brachiaria ramosa (L.) Stapf

Poaceae

Browntop Millet

SNU/CESE/190

29

Brachiaria reptans (L.) C.A.Gardner & C.E.Hubb.

Poaceae

Running Grass

SNU/CESE/085

30

Caesulia axillaris Roxb.

Asteraceae

Pink Node Flower

SNU/CESE/280

31

Cannabis sativa L.

Cannabaceae

Bhang

SNU/CESE/148

32

Capsella bursa-pastoris (L.) Medik.

Brassicaceae

Shepherd’s Purse

SNU/CESE/317

33

Cardamine hirsuta L.

Brassicaceae

Hairy Bitter Cress

SNU/CESE/292

34

Carthamus oxyacantha M.Bieb.

Asteraceae

Wild Safflower

SNU/CESE/434

35

Celosia argentea L.

Amaranthaceae

Safed Murga

SNU/CESE/054

36

Cenchrus ciliaris L.

Poaceae

Buffel Grass

SNU/CESE/346

37

Centaurium pulchellum (Sw.) Druce

Gentianaceae

Pink Centaury

SNU/CESE/349

38

Centella asiatica (L.) Urb.

Apiaceae

Indian Pennywort (Brahmi)

SNU/CESE/383

39

Ceratophyllum demersum L.

Ceratophyllaceae

Coon’s Tail

SNU/CESE/353

40

Chenopodium album L.

Amaranthaceae

Bathua

SNU/CESE/304

41

Chenopodium murale L.

Amaranthaceae

Jangali Bathua

SNU/CESE/416

42

Chloris barbata Sw.

Poaceae

Feather Finger Grass

SNU/CESE/439

43

Chrysopogon zizanioides (L.) Roberty

Poaceae

Khas-khas

SNU/CESE/386

44

Cirsium arvense (L.) Scop.

Asteraceae

Creeping Thistle

SNU/CESE/347

45

Cleome viscosa L.

Cleomaceae

Pili Hurhur

SNU/CESE/030

46

Commelina benghalensis L.

Commelinaceae

Kankawwa

SNU/CESE/231

47

Commelina forskaolii Vahl

Commelinaceae

Kankawwa

SNU/CESE/089

48

Convolvulus prostratus Forssk.

Convolvulaceae

Shankhpushpi

SNU/CESE/343

49

Corchorus aestuans L.

Malvaceae

Chonch

SNU/CESE/077

50

Corchorus olitorius L.

Malvaceae

Pat-Sag

SNU/CESE/384

51

Corchorus tridens L.

Malvaceae

Horn-fruited Jute

SNU/CESE/061

52

Crotalaria medicaginea Lam.

Fabaceae

Trefoil Rattlepod

SNU/CESE/045

53

Crotalaria mysorensis Roth

Fabaceae

Mysore Rattlepod

SNU/CESE/209

54

Croton bonplandianus Baill.

Euphorbiaceae

Ban Tulsi

SNU/CESE/005

55

Cyanotis axillaris (L.) D.Don ex Sweet

Commelinaceae

Kana

SNU/CESE/174

56

Cyanthillium cinereum (L.) H.Rob.

Asteraceae

Sahadevi

SNU/CESE/193

57

Cynodon dactylon (L.) Pers.

Poaceae

Doob

SNU/CESE/006

58

Cyperus alopecuroides Rottb.

Cyperaceae

Foxtail Flatsedge

SNU/CESE/118

59

Cyperus compressus L.

Cyperaceae

Mothi

SNU/CESE/387

60

Cyperus difformis L.

Cyperaceae

Small-Flowered Nutsedge

SNU/CESE/047

61

Cyperus iria L.

Cyperaceae

Rice Flatsedge

SNU/CESE/385

62

Cyperus rotundus L.

Cyperaceae

Motha

SNU/CESE/019

63

Dactyloctenium aegyptium (L.) Willd.

Poaceae

Makra Ghas

SNU/CESE/048

64

Datura innoxia Mill.

Solanaceae

Safed Dhatura

SNU/CESE/007

65

Desmodium gangeticum (L.) DC.

Fabaceae

Shalaparni

SNU/CESE/175

66

Desmodium triflorum (L.) DC.

Fabaceae

Tipatiya

SNU/CESE/218

67

Desmostachya bipinnata (L.) Stapf

Poaceae

Dabh, Kush

SNU/CESE/367

68

Dichanthium annulatum (Forssk.) Stapf

Poaceae

Marvel Grass

SNU/CESE/391

69

Digera muricata (L.) Mart.

Amaranthaceae

Lahsuva

SNU/CESE/368

70

Digitaria ciliaris (Retz.) Koeler

Poaceae

Wild Crabgrass

SNU/CESE/194

71

Dysphania ambrosioides (L.) Mosyakin & Clemants

Amaranthaceae

Sugandha Vastooka

SNU/CESE/096

72

Echinochloa colona (L.) Link

Poaceae

Shama

SNU/CESE/136

73

Echinochloa crus-galli (L.) P.Beauv.

Poaceae

Sanwak

SNU/CESE/371

74

Eclipta prostrata (L.) L.

Asteraceae

Bhringaraj

SNU/CESE/021

75

Eichhornia crassipes (Mart.) Solms

Pontederiaceae

Jal Kumbhi

SNU/CESE/232

76

Eleusine indica (L.) Gaertn.

Poaceae

Jangali Marua

SNU/CESE/112

77

Eragrostis amabilis (L.) Wight & Arn.

Poaceae

Bharbhusi

SNU/CESE/393

78

Eragrostis japonica (Thunb.) Trin.

Poaceae

Pond Lovegrass

SNU/CESE/242

79

Erigeron bonariensis L.

Asteraceae

Flax-leaf Fleabane

SNU/CESE/100

80

Eriochloa procera (Retz.) C.E.Hubb.

Poaceae

Tropical Cupgrass

SNU/CESE/401

81

Euphorbia heterophylla L.

Euphorbiaceae

Lesser Green Poinsettia

SNU/CESE/273

82

Euphorbia hirta L.

Euphorbiaceae

Bari Dudhi, Asthma Weed

SNU/CESE/051

83

Euphorbia hypericifolia L.

Euphorbiaceae

Dudh Mogra

SNU/CESE/196

84

Euphorbia prostrata Aiton

Euphorbiaceae

Choti Dudhi

SNU/CESE/008

85

Euphorbia serpens Kunth

Euphorbiaceae

Dudhi

SNU/CESE/004

86

Evolvulus nummularius (L.) L.

Convolvulaceae

Musakarni

SNU/CESE/394

87

Fimbristylis dichotoma (L.) Vahl

Cyperaceae

Tall Fringe Rush

SNU/CESE/124

88

Fimbristylis ferruginea (L.) Vahl

Cyperaceae

West Indian Fimbry

SNU/CESE/079

89

Fumaria indica (Hausskn.) Pugsley

Papaveraceae

Pittpapra

SNU/CESE/307

90

Gnaphalium purpureum L.

Asteraceae

Purple Cudweed

SNU/CESE/319

91

Gomphrena celosioides Mart.

Amaranthaceae

Prostrate Globe-amaranth

SNU/CESE/062

92

Gonostegia pentandra (Roxb.) Miq.

Urticaceae

Narrow-Leaf Pouzolz's Bush

SNU/CESE/097

93

Grangea maderaspatana (L.) Poir.

Asteraceae

Mustaru

SNU/CESE/358

94

Heliotropium ellipticum Ledeb.

Boraginaceae

Hathisund

SNU/CESE/010

95

Hemarthria compressa (L.f.) R.Br.

Poaceae

Whip Grass

SNU/CESE/125

96

Hydrilla verticillata (L.f.) Royle

Hydrocharitaceae

Hydrilla

SNU/CESE/354

97

Hydrocotyle sibthorpioides Lam.

Araliaceae

Lawn Pennywort

SNU/CESE/126

98

Mesosphaerum suaveolens (L.) Kuntze

Lamiaceae

Vilaiti Tulsi

SNU/CESE/249

99

Imperata cylindrica (L.) Raeusch.

Poaceae

Blady Grass

SNU/CESE/041

100

Indigofera astragalina DC.

Fabaceae

Silky Indigo

SNU/CESE/400

101

Indigofera linifolia (L.f.) Retz.

Fabaceae

Ratnamala

SNU/CESE/250

102

Indigofera tinctoria L.

Fabaceae

Neel

SNU/CESE/149

103

Indigofera tsiangiana Metcalf

Fabaceae

Birdsville Indigo

SNU/CESE/432

104

Justicia japonica Thunb.

Acanthaceae

Common Small Justicia

SNU/CESE/139

105

Lathyrus aphaca L.

Fabaceae

Jangali Matar

SNU/CESE/330

106

Launaea procumbens (Roxb.) Ramayya & Rajagopal

Asteraceae

Peeli Duddhi

SNU/CESE/011

107

Lemna perpusilla Torr.

Araceae

Minute Duckweed

SNU/CESE/411

108

Lepidium didymum L.

Brassicaceae

Jangali Hala

SNU/CESE/302

109

Leptochloa panicea (Retz.) Ohwi

Poaceae

Sprangletop

SNU/CESE/441

110

Leucas cephalotes (Roth) Spreng.

Lamiaceae

Guma

SNU/CESE/197

111

Lindernia ciliata (Colsm.) Pennell

Linderniaceae

Fringed False Pimpernel

SNU/CESE/086

112

Lindernia crustacea (L.) F.Muell.

Linderniaceae

Brittle False Pimpernel

SNU/CESE/087

113

Ludwigia hyssopifolia (G.Don) Exell

Onagraceae

Ban Long

SNU/CESE/198

114

Ludwigia octovalvis (Jacq.) P.H.Raven

Onagraceae

Ban Long

SNU/CESE/226

115

Lysimachia loeflingii F.J.Jiménez & M.Talavera

Primulaceae

Krishnaneel

SNU/CESE/106

116

Malva parviflora L.

Malvaceae

Panirak

SNU/CESE/320

117

Malvastrum coromandelianum (L.) Garcke

Malvaceae

Kharenti

SNU/CESE/018

118

Mazus pumilus (Burm.f.) Steenis

Phrymaceae

Japanese Mazus

SNU/CESE/023

119

Mecardonia procumbens (Mill.) Small

Plantaginaceae

Baby Jump Up

SNU/CESE/083

120

Medicago monantha (C.A.Mey.) Trautv.

Fabaceae

Single-Flowered Medick

SNU/CESE/335

121

Medicago lupulina L.

Fabaceae

Black Medic

SNU/CESE/294

122

Medicago polymorpha L.

Fabaceae

Toothed Medic

SNU/CESE/424

123

Melilotus albus Medik.

Fabaceae

Safed Ban Methi

SNU/CESE/337

124

Melilotus indicus (L.) All.

Fabaceae

Ban Methi

SNU/CESE/287

125

Melochia corchorifolia L.

Malvaceae

Chitrabeej

SNU/CESE/050

126

Mollugo nudicaulis Lam.

Molluginaceae

Naked-stem Carpetweed

SNU/CESE/164

127

Murdannia nudiflora (L.) Brenan

Commelinaceae

Kansura

SNU/CESE/177

128

Nepeta hindostana (B.Heyne ex Roth) Haines

Lamiaceae

Billi Lotan

SNU/CESE/200

129

Nicotiana plumbaginifolia Viv.

Solanaceae

Jangali Tambakoo

SNU/CESE/012

130

Nymphaea pubescens Willd.

Nymphaeaceae

Kumud

SNU/CESE/276

131

Nymphoides cristata (Roxb.) Kuntze

Menyanthaceae

Kumudini

SNU/CESE/275

132

Oldenlandia corymbosa L.

Rubiaceae

Daman Pappar

SNU/CESE/013

133

Oxalis corniculata L.

Oxalidaceae

Amrul, Khatti-buti

SNU/CESE/084

134

Oxalis debilis var. corymbosa (DC.) Lourteig

Oxalidaceae

Khatti-mithi

SNU/CESE/338

135

Panicum antidotale Retz.

Poaceae

Blue Panic Grass

SNU/CESE/405

136

Parthenium hysterophorus L.

Asteraceae

Gajar Ghas

SNU/CESE/014

137

Paspalum distichum L.

Poaceae

Knotgrass

SNU/CESE/440

138

Paspalum scrobiculatum L.

Poaceae

Kodo

SNU/CESE/063

139

Dicliptera paniculata (Forssk.) I.Darbysh.

Acanthaceae

Atrilal

SNU/CESE/267

140

Persicaria barbata (L.) H.Hara

Polygonaceae

Bearded Knotweed

SNU/CESE/398

141

Persicaria lanigera (R.Br.) Soják

Polygonaceae

Pink Knotweed

SNU/CESE/024

142

Phalaris minor Retz.

Poaceae

Mandusi

SNU/CESE/326

143

Phragmites karka (Retz.) Trin. ex Steud.

Poaceae

Narkul

SNU/CESE/406

144

Phyla nodiflora (L.) Greene

Verbenaceae

Jal Buti

SNU/CESE/070

145

Phyllanthus amarus Schumach. & Thonn.

Phyllanthaceae

Bhuiaonla

SNU/CESE/132

146

Phyllanthus maderaspatensis L.

Phyllanthaceae

Hajarmani

SNU/CESE/043

147

Phyllanthus tenellus Roxb.

Phyllanthaceae

Mascarene Island Leaf-flower

SNU/CESE/437

148

Physalis angulata L.

Solanaceae

Rasbhari

SNU/CESE/037

149

Physalis peruviana L.

Solanaceae

Rasbhari

SNU/CESE/410

150

Pistia stratiotes L.

Araceae

Water Lettuce

SNU/CESE/395

151

Pluchea lanceolata (DC.) C.B.Clarke

Asteraceae

Rasna

SNU/CESE/064

152

Poa annua L.

Poaceae

Annual Bluegrass

SNU/CESE/421

153

Polygonum plebeium R.Br.

Polygonaceae

Small Knotweed

SNU/CESE/308

154

Portulaca oleracea L.

Portulacaceae

Kulpha, Noni

SNU/CESE/351

155

Portulaca quadrifida L.

Portulacaceae

Chicken Weed

SNU/CESE/381

156

Potamogeton crispus L.

Potamogetonaceae

Curly-Leaf Pondweed

SNU/CESE/412

157

Ranunculus sceleratus L.

Ranunculaceae

Jaldhaniya

SNU/CESE/352

158

Rorippa palustris (L.) Besser

Brassicaceae

Chamsuru

SNU/CESE/315

159

Rumex dentatus L.

Polygonaceae

Jangali Palak

SNU/CESE/419

160

Rumex hypogaeus T.M.Schust. & Reveal

Polygonaceae

Three Corner Jack

SNU/CESE/314

161

Rungia pectinata (L.) Nees

Acanthaceae

Comb Rungia

SNU/CESE/413

162

Saccharum bengalense Retz.

Poaceae

Munj, Sarkanda

SNU/CESE/404

163

Saccharum spontaneum L.

Poaceae

Kaans

SNU/CESE/379

164

Sagittaria guayanensis Kunth

Alismataceae

Guyanese Arrowhead

SNU/CESE/199

165

Scoparia dulcis L.

Plantaginaceae

Ghoda Tulsi

SNU/CESE/001

166

Senna obtusifolia (L.) H.S.Irwin & Barneby

Fabaceae

Chakwar

SNU/CESE/147

167

Senna occidentalis (L.) Link

Fabaceae

Kasundi

SNU/CESE/016

168

Sesamum indicum L.

Pedaliaceae

Safed Til

SNU/CESE/184

169

Sesbania bispinosa (Jacq.) W.Wight

Fabaceae

Dhaincha

SNU/CESE/065

170

Setaria pumila (Poir.) Roem. & Schult.

Poaceae

Bandra

SNU/CESE/128

171

Setaria verticillata (L.) P.Beauv.

Poaceae

Laptuna

SNU/CESE/374

172

Sida acuta Burm.f.

Malvaceae

Baraira

SNU/CESE/068

173

Sida cordata (Burm.f.) Borss.Waalk.

Malvaceae

Kharenti

SNU/CESE/203

174

Sida cordifolia L.

Malvaceae

Bala

SNU/CESE/215

175

Silene conoidea L.

Caryophyllaceae

Cone Catchfly

SNU/CESE/310

176

Sisymbrium irio L.

Brassicaceae

Khubkalan

SNU/CESE/322

177

Solanum americanum Mill.

Solanaceae

Makoy

SNU/CESE/286

178

Solanum surattense Burm. f.

Solanaceae

Kateli

SNU/CESE/032

179

Sonchus asper (L.) Hill

Asteraceae

Dudhi

SNU/CESE/311

180

Sonchus brachyotus DC.

Asteraceae

Dudhali

SNU/CESE/117

181

Sorghum halepense (L.) Pers.

Poaceae

Jangali Jowar

SNU/CESE/205

182

Spergula arvensis L.

Caryophyllaceae

Ban Dhania

SNU/CESE/425

183

Spermacoce articularis L.f.

Rubiaceae

Guthari

SNU/CESE/058

184

Sphenoclea zeylanica Gaertn.

Sphenocleaceae

Goose Weed

SNU/CESE/204

185

Spirodela polyrhiza (L.) Schleid.

Araceae

Common Duck Meat

SNU/CESE/414

186

Stellaria media (L.) Vill.

Caryophyllaceae

Buch-Bucha

SNU/CESE/296

187

Stuckenia pectinata (L.) Börner

Potamogetonaceae

Sago Pondweed

SNU/CESE/433

188

Symphyotrichum squamatum (Spreng.) G.L.Nesom

Asteraceae

Annual Saltmarsh Aster

SNU/CESE/456

189

Tephrosia pumila (Lam.) Pers.

Fabaceae

Indigo Sauvage

SNU/CESE/380

190

Tephrosia purpurea (L.) Pers.

Fabaceae

Sharpunkha

SNU/CESE/069

191

Tephrosia villosa (L.) Pers.

Fabaceae

Hoary Tephrosia

SNU/CESE/388

192

Trianthema portulacastrum L.

Aizoaceae

Vishakhapara

SNU/CESE/017

193

Tribulus terrestris L.

Zygophyllaceae

Gokhru

SNU/CESE/442

194

Tridax procumbens (L.) L.

Asteraceae

Ghamra

SNU/CESE/099

195

Trifolium tomentosum L.

Fabaceae

Tipatiya Ghaas

SNU/CESE/283

196

Triumfetta rhomboidea Jacq.

Malvaceae

Chinese Bur

SNU/CESE/234

197

Typha domingensis Pers.

Typhaceae

Patera

SNU/CESE/390

198

Utricularia stellaris L.f.

Lentibulariaceae

Bladderwort

SNU/CESE/408

199

Vallisneria spiralis L.

Hydrocharitaceae

Tape Grass

SNU/CESE/431

200

Verbascum chinense (L.) Santapau

Scrophulariaceae

Gadar Tambaku

SNU/CESE/144

201

Verbascum thapsus L.

Scrophulariaceae

Jangali Tambaku

SNU/CESE/134

202

Veronica agrestis L.

Plantaginaceae

Field Speedwell

SNU/CESE/288

203

Veronica anagallis-aquatica L.

Plantaginaceae

Water Speedwell

SNU/CESE/324

204

Vicia sativa L.

Fabaceae

Matri

SNU/CESE/298

205

Xanthium strumarium L.

Asteraceae

Sankhahuli

SNU/CESE/257

 

Shrubs

 

 

 

206

Abutilon indicum (L.) Sweet

Malvaceae

Kanghi

SNU/CESE/157

207

Calotropis gigantea (L.) Dryand.

Apocynaceae

Safed Aak

SNU/CESE/365

208

Calotropis procera (Aiton) Dryand.

Apocynaceae

Aak, Madar

SNU/CESE/026

209

Capparis sepiaria L.

Capparaceae

Wild Caper Bush

SNU/CESE/102

210

Clerodendrum phlomidis L.f.

Verbenaceae

Arni

SNU/CESE/247

211

Ficus palmata Forssk.

Moraceae

Jangali Anjir

SNU/CESE/115

212

Ipomoea carnea Jacq.

Convolvulaceae

Behaya

SNU/CESE/271

213

Jatropha gossypiifolia L.

Euphorbiaceae

Bellyache Bush

SNU/CESE/436

214

Justicia adhatoda L.

Acanthaceae

Arus, Arusa

SNU/CESE/333

215

Lantana camara L.

Verbenaceae

Raimuniya

SNU/CESE/080

216

Mimosa rubicaulis Lam.

Fabaceae

Himalayan Mimosa

SNU/CESE/207

217

Phyllanthus reticulatus Poir.

Phyllanthaceae

Kale Madhu Ka Per

SNU/CESE/116

218

Ricinus communis L.

Euphorbiaceae

Arandi

SNU/CESE/213

219

Solanum torvum Sw.

Solanaceae

Bhankatiya

SNU/CESE/142

220

Tamarix indica Willd.

Tamaricaceae

Jhau

SNU/CESE/396

221

Urena lobata L.

Malvaceae

Bachita, Lapetua

SNU/CESE/171

222

Withania somnifera (L.) Dunal

Solanaceae

Ashwagandha

SNU/CESE/145

223

Ziziphus nummularia (Burm.f.)Wight & Arn.

Rhamnaceae

Jhar Beri

SNU/CESE/240

 

Climbers/Creepers

 

 

 

224

Abrus precatorius L.

Fabaceae

Ratti

SNU/CESE/382

225

Cajanus scarabaeoides (L.) Thouars

Fabaceae

Showy Pigeonpea

SNU/CESE/222

226

Cayratia trifolia (L.) Domin

Vitaceae

Three-Leaved Wild Vine

SNU/CESE/135

227

Cissampelos pareira L.

Menispermaceae

Patha

SNU/CESE/191

228

Citrullus colocynthis (L.) Schrad.

Cucurbitaceae

Colocynth

SNU/CESE/407

229

Coccinia grandis (L.) Voigt

Cucurbitaceae

Kundru

SNU/CESE/109

230

Convolvulus arvensis L.

Convolvulaceae

Hirankhuri

SNU/CESE/305

231

Cucumis melo L.

Cucurbitaceae

Kachariya

SNU/CESE/039

232

Cuscuta chinensis Lam.

Convolvulaceae

Amar Bel

SNU/CESE/034

233

Cuscuta reflexa Roxb.

Convolvulaceae

Amar Bel

SNU/CESE/301

234

Ipomoea aquatica Forssk.

Convolvulaceae

Kalmi Sag

SNU/CESE/137

235

Ipomoea coptica (L.) Roth ex Roem. & Schult.

Convolvulaceae

Egyptian Morning Glory

SNU/CESE/091

236

Ipomoea eriocarpa R. Br.

Convolvulaceae

Tiny Morning Glory

SNU/CESE/210

237

Ipomoea obscura (L.) Ker Gawl.

Convolvulaceae

Obscure Morning Glory

SNU/CESE/131

238

Ipomoea pes-tigridis L.

Convolvulaceae

Tiger Foot Morning Glory

SNU/CESE/163

239

Ipomoea triloba L.

Convolvulaceae

Three-lobe Morning Glory

SNU/CESE/252

240

Merremia hederacea (Burm. f.) Hallier f.

Convolvulaceae

Ivy Woodrose

SNU/CESE/244

241

Mukia maderaspatana (L.) M.Roem.

Cucurbitaceae

Bilari

SNU/CESE/127

242

Operculina turpethum (L.) Silva Manso

Convolvulaceae

Nisoth, Pithori

SNU/CESE/245

243

Oxystelma esculentum (L. f.) Sm.

Apocynaceae

Dudhialata

SNU/CESE/166

244

Rhynchosia capitata (Roth) DC.

Fabaceae

Kulata, Kulthi

SNU/CESE/211

245

Rhynchosia minima (L.) DC.

Fabaceae

Kulata, Kulthi

SNU/CESE/212

246

Teramnus labialis (L.f.) Spreng.

Fabaceae

Van Udad

SNU/CESE/217

247

Trichosanthes cucumerina L.

Cucurbitaceae

Jangali Chachinda

SNU/CESE/377

 

Trees

 

 

 

248

Acacia nilotica (L.) Delile

Fabaceae

Babool

SNU/CESE/361

249

Albizia lebbeck (L.) Benth.

Fabaceae

Siris

SNU/CESE/289

250

Azadirachta indica A.Juss.

Meliaceae

Neem

SNU/CESE/348

251

Butea monosperma (Lam.) Taub.

Fabaceae

Dhak, Palash

SNU/CESE/279

252

Cordia dichotoma G.Forst.

Boraginaceae

Lasoda

SNU/CESE/003

253

Crateva adansonii ssp. odora (Buch.-Ham.) Jacobs

Capparaceae

Barna, Barni

SNU/CESE/095

254

Dalbergia sissoo DC.

Fabaceae

Shisham

SNU/CESE/418

255

Diospyros montana Roxb.

Ebenaceae

Bistendu

SNU/CESE/357

256

Ficus religiosa L.

Moraceae

Peepal

SNU/CESE/031

257

Holoptelea integrifolia Planch.

Ulmaceae

Chilbil

SNU/CESE/438

258

Leucaena leucocephala (Lam.) de Wit

Fabaceae

Subabul

SNU/CESE/256

259

Melia azedarach L.

Meliaceae

Bakain

SNU/CESE/098

260

Mitragyna parvifolia (Roxb.) Korth.

Rubiaceae

Kaim

SNU/CESE/092

261

Morus alba L.

Moraceae

Shahtoot

SNU/CESE/042

262

Phoenix sylvestris (L.) Roxb.

Arecaceae

Jangali Khajoor

SNU/CESE/201

263

Prosopis cineraria (L.) Druce

Fabaceae

Khejri

SNU/CESE/282

264

Prosopis juliflora (Sw.) DC.

Fabaceae

Vilayati Babul

SNU/CESE/180

265

Salix tetrasperma Roxb.

Salicaceae

Bed-laila

SNU/CESE/290

266

Sesbania sesban (L.) Merr.

Fabaceae

Rawasan

SNU/CESE/066

267

Streblus asper Lour.

Moraceae

Sihora

SNU/CESE/430

268

Syzygium cumini (L.) Skeels

Myrtaceae

Jamun

SNU/CESE/360

269

Ziziphus jujuba Mill.

Rhamnaceae

Ber

SNU/CESE/246

 

Lycophytes and Fern

 

 

 

270

Ampelopteris prolifera (Retz.) Copel.

Thelypteridaceae

Walking Fern

SNU/CESE/094

271

Equisetum ramosissimum Desf.

Equisetaceae

Branched Horsetail

SNU/CESE/113

272

Marsilea quadrifolia L.

Marsileaceae

Chaupatiya

SNU/CESE/140

 

 

Table 2. Floristic details of studies undertaken in the National Capital Territory Delhi (NCT Delhi), and National Capital Region (NCR) under western Uttar Pradesh.

Study area

Floristic study

Total plant species recorded

No. of plant

families

No. of

genera

Reference

 

Chithara, NCR (7.7 km2)

272

69

204

Present study

NCT Delhi (1,483 km2)

531

-

-

Maheshwari (1963)

Baghpat (NCR)

566

102

371

Ahamed & Gupta (2010)

Bulandshahr (NCR)

376

-

-

Singh (1969)

Ghaziabad (NCR)

1,654*

171

868

Vardhana (2007)

Meerut (NCR)

862

118

521

Shishodia (2013)

Muzaffarnagar (NCR)

577

77

321

Malik (2015)

*included wild, cultivated, and non-native species.
All floristic details include non-native and invasive species along with native species to the region.

 

For figures & images - - click here for full PDF

 

 

REFERENCES

 

Ahamed, N. & A.K. Gupta (2010). An analysis of flora of Baghpat district in Uttar Pradesh, India. Indian Journal of Forestry 33(3): 405–418. https://doi.org/10.54207/bsmps1000-2010-1KR94K

Duthie, J.F. (1960). Flora of the Upper Gangetic Plain and of the Adjacent Siwalik and Sub-Himalayan Tracts. Reprint edition. Botanical Survey of India, Calcutta, 554 pp.

Maheshwari, J.K. (1963). The Flora of Delhi. Council of Scientific and Industrial Research, New Delhi, 447 pp.

Maheshwari, J.K. (1966). Illustrations of the Flora of Delhi. CSIR, New Delhi, India, 282 pp.

Malik, V. (2015). Herbaceous flora of Muzaffarnagar district (U.P.). Scholars Academic Journal of Biosciences 3(2B): 182–196.

POWO (2024). Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. https://powo.science.kew.org/. Accessed 07 October 2024.

Sen, S. & R. Chakraborty (2017). Revival, modernization and integration of Indian traditional herbal medicine in clinical practice: importance, challenges and future. Journal of Traditional and Complementary Medicine 7(2): 234–244. https://doi.org/10.1016/j.jtcme.2016.05.006

Sharma, A.K. & J.S. Dhakre (1995). Flora of Agra District. Botanical Survey of India, Calcutta, India, 356 pp.

Sharma, J.K., A.K. Tripathi & M. Ahmad (2018). Illustrated Flora: Part of Western Uttar Pradesh and Delhi NCR, India. Siya Publishing House and Shiv Nadar University, 606 pp.

Shishodia, J. (2013). Study of flora of district Meerut in form of digital herbarium database. Ph.D. Thesis. Department of Botany, Chaudhary Charan Singh University, Meerut, Uttar Pradesh, India, 632 pp.

Singh, N.P. (1969). Flora of Bulandshahr district. Nelumbo 11(1–2): 1–22. https://doi.org/10.20324/nelumbo/v11/1969/75822

Tripathi, A.K. & J.K. Sharma (2019). Occurrence of Symphyotrichum squamatum (Spreng.) G. L. Nesom in Uttar Pradesh, India: a new record. Modern Phytomorphology 13: 26–29. https://doi.org/10.5281/zenodo.3518822

Tripathi, A.K., J.K. Sharma & M. Ahmad (2018). Emex australis Steinh. — a new record for Uttar Pradesh and upper Gangetic Plains of India. Indian Journal of Plant Sciences 7(2): 1–5.

Tripathi, A.K., J.K. Sharma & M. Ahmad (2020). Traditional Usages and Chemical Constituents of High Value Medicinal Plants Recorded from Chithara Village Panchayat (Gautam Buddha Nagar) of Uttar Pradesh, India, pp. 221–273. In: Singh, B. & Y.P. Sharma (eds.). Plants for Novel Drug Molecules: Ethnobotany to Ethnopharmacology. New India Publishing Agency, New Delhi, 590 pp.

Tripathi, A.K., J.K. Sharma, G.D. Bhatt & M. Ahmad (2019). Gregarious colonization of Tribulus terrestris L. (Puncture Vine): ecological implications. Environment and Ecology 37(3B): 1092–1099.

Vardhana, R. (2007). Flora of Ghaziabad District. Shree Publishers & Distributors, New Delhi, 639 pp.