Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 July 2024 | 16(7): 25604–25608
ISSN 0974-7907
(Online) | ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.8859.16.7.25604-25608
#8859 | Received 28 November 2023 | Final received 29 June 2024 | Finally
accepted 03 July 2024
The opportunistic feeding behaviour of Schistura notostigma (Teleostei: Nemacheilidae) in tropical mountain streams in Sri Lanka
J. Bandara
1, M.P. Gunawardena 2 &
R.T.P Jayasuriya 3
1,2,3 Faculty of Science, Horizon
Campus, Knowledge City Malabe, KCM Drive, Off Millennium Road, Malabe, 10115,
Sri Lanka.
2 Biodiversity Educational Research
Initiative (BERI), LE CUBE, No. 130, High Level Road, Colombo 06, 00600, Sri
Lanka.
2 Thema Collection, LE CUBE, No. 130,
High Level Road, Colombo 06, 00600, Sri Lanka.
1 janaminabandara@gmail.com
(corresponding author), 2 medhisha@gmail.com, 3 ravishkajayasuriya@gmail.com
Editor: Rohan Pethiyagoda,
Australian Museum, Sydney, Australia. Date of publication: 26 July 2024
(online & print)
Citation: Bandara, J., M.P. Gunawardena & R.T.P Jayasuriya (2024). The opportunistic feeding
behaviour of Schistura notostigma
(Teleostei: Nemacheilidae)
in tropical mountain streams in Sri Lanka. Journal of Threatened Taxa 16(7): 25604–25608. https://doi.org/10.11609/jott.8859.16.7.25604-25608
Copyright: © Bandara et al. 2024. Creative Commons Attribution 4.0
International License. JoTT allows unrestricted use, reproduction, and
distribution of this article in any medium by providing adequate credit to the
author(s) and the source of publication.
Funding: This research was supported by funding from BERI and Thema Collection, Sri Lanka.
Competing interests: The authors declare no competing interests.
Acknowledgements: The authors are grateful to Rohan Pethiyagoda for reviewing an earlier version
of the manuscript draft, and to Hiranya Sudasinghe for providing constructive
comments and reviewing the current version of the manuscript. The authors also
thank Madura De Silva for permitting the use of his photographs in this paper.
Abstract: Schistura notostigma is a bottom-dwelling freshwater
fish endemic to Sri Lanka. In this study, we document an opportunistic
feeding behavior exhibited by this loach species in close association
with Garra ceylonensis
and Plesiopuntius bimaculatus
in the streams of high-elevation headwater regions of the Knuckles
Mountain Range and Piduruthalagala Mountain Range in
Sri Lanka.
Keywords: Feeding ecology, behavioural ecology, coexistence, commensalism
Schistura notostigma (Bleeker, 1863) is a small,
bottom-dwelling freshwater fish endemic to Sri Lanka. It features an elongated,
moderately cylindrical body that is ventrally flattened and slender, and
possesses three pairs of small barbels, which
characteristics are shared by many benthic freshwater fishes (Pethiyagoda 1991; Sundarabarathy
et al. 2001; Herath 2009; Shirantha
2021). In this study, we document instances of opportunistic feeding behavior
exhibited by this loach. These behaviors were observed frequently in the close
association with two species of the family Cyprinidae,
Garra ceylonensis
(Bleeker, 1863) and Plesiopuntius bimaculatus (Bleeker, 1863), in streams situated in
high elevation headwater regions of the Knuckles Mountain Range (KMR) and Piduruthalagala Mountain Range (PMR).
Schistura notostigma (Image 1A) is one of the few
freshwater fishes found at elevations exceeding 1,000 m in Sri Lanka, along
with Garra ceylonensis
(Image 01B), Plesiopuntius bimaculatus (Image 01C) and Devario
spp. (Pethiyagoda & Sudasinghe
2021). Although S. notostigma can
sometimes be found at elevations above 1,300 m, the other species mentioned
generally inhabit elevations up to 1,000 m, albeit with rare exceptions (Pethiyagoda & Sudasinghe
2021; Sudasinghe et al. 2021, 2023). Despite our
observations indicating that S. notostigma
could occasionally be the sole freshwater fish in high-elevation habitats,
however, it exhibits a notable inclination to associate closely with G. ceylonensis when present in the ichthyo-community as
associates.
Materials and Methods
The present study was conducted
as part of an ongoing research project on Sri Lankan freshwater fishes. The
study was carried out in Mandaramnuwara, situated on
the eastern slopes of PMR at an elevation of 1,000 m. PMR study site is a
circular area with a 5-km radius centered on coordinates 7.06380N,
80.76520E, situated in the Belihul Oya (river) headwaters, near Kalu Palama
Ella, and in Medadumbara, on the southern flank of
KMR at 590 m. KMR study site is a circular area of 5-km radius, centered at
7.34750N, 80.82020E in the Heel Oya
and Galmal Oya headwater
regions (Figure 1). Five points were arbitrarily selected for observation from
each site within the specified range. These observations spanned a 12-month
period from November 2022 to October 2023. Observations were made monthly (five
days per month from 0700–1800 h) using remote underwater video footage (GoPro
Hero 9 action camera) and snorkelling. Fish
identification was performed in the field using the latest field guides (De
Silva et al. 2015; Shirantha 2021) and later
confirmed by reference to recent publications (Sudasinghe
2017, 2018). During the species identification process, fishes were captured
using a scoop-net and immediately released to their original habitats.
Observations and
Discussion
Garra ceylonensis and Schistura
notostigma (at an elevation range of 590–1,350 m
in KMR and PMR) exhibit a close association and a commensalism relationship as
follows: G. ceylonensis was observed
grazing on algae on the rocky substrate, using its sucker mouth (Sudasinghe et al. 2021), causing small invertebrates and
other food particles to be detached. The opportunistic species S. notostigma then exploits this disturbance by
browsing on the dislodged food particles (see Image 2A,B).
Analysis of diets,
notwithstanding different degrees of selectivity for various food categories,
showed both species to be omnivorous (confirmed through gut content analysis in
prior studies and references in: Moyle & Senanayake
1984; Wikramanayake & Moyle 1989; Sundarabarathy et al. 2001, 2005; Shirantha
2004; Pethiyagoda & Sudasinghe
2021). However, S. notostigma was
found to primarily feed on a variety of small invertebrates, especially
caddisfly larvae (Trichoptera: Hydropsychidae
and Rhyacophilidae) and small mayfly
larvae (Ephemeroptera: Leptophlebeidae) (Moyle & Senanayake 1984). Furthermore, Moyle & Senanayake (1984) suggest carnivory as the dominant mode of
feeding for this loach, highlighting the short intestine (less than 1 × body
length) and the observed willingness of S. notostigma
to attack small benthic prey. On the other hand, G. ceylonensis is mentioned as a benthic algae feeder
in Sudasinghe (2021) by referring to Lujan &
Conway (2015) and Kottelat (2020). Moreover, Moyle
& Senanayake (1984) also showed G. ceylonensis to be primarily a detritus and diatom
feeder, based on a gut content analysis in Costa & Fernando (1967).
Furthermore, studies conducted by Wikramanayake &
Moyle (1989) and Moyle & Senanayake (1984) in
rainforest streams and lowland wet zone streams show that there is no
statistically significant dietary overlap among G. ceylonensis
and S. notostigma. Therefore, based on
the diet and behaviour, we hypothesize this
association as a commensal relationship, in which S. notostigma
benefits substantially more, rather than a mutualism in which both species
benefit.
A similar set of observations
were also recorded with schools of Plesiopuntius
bimaculatus in the KMR during the study period
(recorded in five occasions specifically during March 2023 and April 2023),
which also confirmed the opportunistic feeding behaviour
of S. notostigma. Here, S. notostigma followed the schools of foraging Pl.
bimaculatus, feeding on food particles sinking to
the substrate.
In this case
of S. notostigma, given the absence of
any recorded negative interactions up to date, whether of a predatory or
competitive nature between these species (G. ceylonensis
and S. notostigma or Pl.
bimaculatus and S. notostigma), we suggest that opportunistic feeding behaviours and close associations may enable these species
to access food sources more easily and with less competition. This becomes
especially pertinent in high-elevation tropical mountain regions, where
co-evolved fishes and other vertebrate species are recognized to have a high
degree of adaptation to their environments, as described in Abell et al.
(2008).
In addition
to S. notostigma, similar opportunistic
feeding behaviours have been observed in other
benthic loach species, including Misgurnus anguillicaudatus (Cantor, 1842) as documented by VanHaitsma (2020) and McNair et al. (2023), Misgurnus fossilis
(Linnaeus, 1758) as reported by Pyrzanowski et al.
(2019), and Barbatula barbatula
(Linnaeus, 1758) as noted by Andrei et al. (2015) and Worischka
et al. (2015). Furthermore, Rice et al. (2019) too, make
reference to several unique benthic feeding behaviours
of fishes.
Pethiyagoda & Sudasinghe
(2021) also make reference to a similar set of
ecological interactions among Sri Lankan freshwater fishes, originally
documented by Schut et al. (1984). These interactions
involve Rohanella titteya
(Deraniyagala, 1929) and Pl. bimaculatus, as well as juvenile Dawkinsia
fillementosa (Valenciennes, 1844) and adult Pethia cumingii
(Günther, 1868). These associations can be succinctly summarized as
optimizations of dietary habits and the avoidance of shared predators.
Apart from G.
ceylonensis, S. notostigma,
and Pl. bimaculatus, only Devario malabaricus
(Jerdon, 1849) and Poecilia
reticulata (Peters, 1859) were found inhabiting the high-elevation regions
in KMR (600 m) (Pethiyagoda & Sudasinghe
2021; Bandara & Gunawardena 2023a,b). No other freshwater fishes were recorded in headwaters
of the PMR study site, except for G. ceylonensis
and S. notostigma. Generally, it
shows a lower fish diversity in high-elevation tropical freshwater fish
assemblages recorded above 1,000 m elevation, when compared to lowland
assemblages in the island’s south west wet zone (Sumith
et al. 2011; Pethiyagoda & Sudasinghe
2021; Sudasinghe et al. 2023). In this case we
believe that the opportunistic feeding of S. notostigma
reported here could be a common behaviour with the
co-occurring benthic species G. ceylonensis
and Pl. bimaculatus.
Conclusion
In
conclusion, frequent occurrences of the aforementioned ecological associations
and interactions provide insight into the feeding behaviours
of this benthic loach species. Moreover, these records also offer vital
information that can be used to understand the ecological structure of tropical
freshwater fish assemblages in high-elevation mountain streams (1,000 m) in Sri
Lanka.
For figure
& images - - click here for full PDF
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