Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 November 2023 | 15(11): 24280–24283
ISSN 0974-7907
(Online) | ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.8630.15.11.24280-24283
#8630 | Received 11 July 2023 | Final received 21 August 2023 | Finally
accepted 13 October 2023
A preliminary observation on the
nesting of the Indochinese Roller Coracias affinis Horsfield, 1840
(Aves: Coraciiformes: Coraciidae)
in Assam and northern West Bengal, India
Sachin Ranade 1, Jay Gore
2 & Sonali Ranade 3
1,2 Vulture Conservation Breeding
Centre, Rani, Kamrup, Assam 781131, India.
3 Department of Forest Genetics and
Plant Physiology, Umeå Plant Science Centre, Swedish
University of Agricultural Sciences, SE, Umeå.
1 s.ranade@bnhs.org (corresponding
author), 2 jaygore54@gmail.com, 3 sonalideo@yahoo.com
Editor: H. Byju,
Coimbatore, Tamil Nadu, India. Date of publication: 26 November
2023 (online & print)
Citation: Ranade, S., J. Gore & S. Ranade (2023). A preliminary
observation on the nesting of the Indochinese Roller Coracias
affinis Horsfield, 1840
(Aves: Coraciiformes: Coraciidae)
in Assam and northern West Bengal, India. Journal of Threatened Taxa 15(11): 24280–24283. https://doi.org/10.11609/jott.8630.15.11.24280-24283
Copyright: © Ranade et al. 2023. Creative Commons Attribution 4.0 International
License. JoTT
allows unrestricted use, reproduction, and distribution of this article in any
medium by providing adequate credit to the author(s) and the source of
publication.
Funding: None.
Competing interests: The authors declare no competing interests.
Acknowledgements: We thank the Bombay Natural
History Society and forest departments of Assam and West Bengal for constant
support and encouragement.
Abstract: The nesting of Indochinese
Roller was observed in Rani, Kamrup, Assam for 49
days in April–May 2022. The male was noted hunting more frequently compared to
the female, but its prey items consisted of smaller biomass than those the
female hunted. Opportunistically, more nests were recorded at Buxa Tiger Reserve, West Bengal and Kaziranga
National Park, Assam. The preferred tree species for nesting were Sal Shorea robusta,
Mynah Tetrameles nudiflora,
and Koroi Albizia
procera; the preferred nest height was 9.2
m.
Keywords: Buxa,
cavity nesting, Kaziranga, parental role, prey base.
The Indochinese Roller Coracias affinis Horsfield, 1840 also known as Black-billed Roller and
Burmese Roller, has been recently raised to the species level (Johansson
et al. 2018; Gill et al. 2021). Formerly
it was considered the subspecies of Indian Roller Coracias
benghalensis. Its ‘unstreaked
smoky-purplish brown face and breast’ differentiate it from the Indian Roller C.
benghalensis. Its distribution is across Nepal, Bhutan, northeastern
India to south-central China, northern Malay Peninsula, and Indochina (Ali
& Ripley 1987).
The nesting of Indochinese Roller
was observed at Belguri Village (25.99960N,
91.54840E), Rani, Assam since 2007 (n = 16). Each year, a pair
occupied the same nest in the Sal Shorea robusta although we could not ensure it was the pair of
the same individuals. In 2022, we followed the focal animal sampling method and
recorded the parents’ and nestlings’ activities once per minute (Altmann 1974;
Palmer et al. 2001). The observations were categorised
in three sloughs 0600–1000 h (n = 1,417), 1000–1400 h (n = 1,444) and 1400–1800
h (n = 2,151). The activities were compared between these sloughs and
significance was checked with chi square test. The observations period was from
13 April–31 May 2022 when two nestlings fledged out (Image 1). The young ones
followed the parents for about 90 days during which the begging calls for food
were noted (https://xeno-canto.org/819166). We also made opportunistic observation
on the nesting of the species in Buxa Tiger Reserve
(26.67440N, 89.74720E), Alipurduar
District, West Bengal and in Kaziranga National Park
(26.57580N, 93.16700E), Golaghat
District, Assam.
The height and girth of the
nesting tree Sal was 26.8 m and 2.3 m, respectively, while the nest was at a
height of 11.6 m. The parents perched (n = 113) around the nest during the
breeding period on 10 trees which were at a distance of 5.2 m to 34.4 m from
the nesting tree. Most of the trees were young trees including Tectona grandis
(8), one mature Shorea robusta,
and one mature Moringa oleifera.
The tree heights ranged 7.6–27 m and the most frequent perching height
was around 11.6 m on all of these trees.
The prey items were photographed
and identified with keys and opportunistically similar items were weighed with
digital weighing balance to get an idea of the biomass.
Results
Observations of the nesting pair
The male and female have distinct
characteristics as follows: females are slightly larger
and males possess brilliant blue colour on the throat
(Image 2 & 3). The incubating and brooding parents could not be observed,
but the activities of the parents were recorded for perching, hunting and
carrying the food.
Overall, the male was engaged in
hunting for a longer duration throughout the day as compared to the female,
which was statistically significant (P <0.0001). The hunting time of the
male and the female was recorded for all the time slots throughout the day
(0600 h–1000 h, X2
= 38.534, df = 1, P < 0.0001; 1000–1400 h, X2 = 15.244, df = 1, P <0.0001; 1400h–1800 h, X2 = 15.254, df = 1, P <0.0001. Female was found to be perched for
significantly (P <0.0001) higher number of times as compared to the male,
except for the time slot between 1400 h to 1800 h where both male and female
were observed to be perched equally (0600–1000 h, X2 = 40.998, df = 1,
P <0.0001; 1000–1400 h, X2
= 46.815, df = 1, P <0.0001; 1400–1800 h, X2= 0.252, df = 1, P = 0.6154) (Figure 1).
The prey base
In all 100 successful hunting and
food carrying attempts that were recorded, the male captured 56 prey items and
the female captured 44 prey items (Table 1); but, with reference to the
biomass, the female captured more biomass than the male (the prey captured by
the female was about 100.6 g while in the case of the male, it was 29.1 g). The
female hunted larger prey including mainly frogs (Fejervarya spp.),
snakes (Dendrelaphis spp.), and a
lizard (Calotes spp.). The male captured more
insects, and the cicadas were the most common among the identified insects (Haphsa scitula and Dundubia annandalei)
(Anonymous 2022a,b). Out of the 86
insect prey captured by the pair, 31% were identified while 69% of the
prey captures were unknown. The identified prey included cicada (20%),
cockroach (1%), dragonfly (5%), mantis (2%), grasshopper (1%,) and termite (alate) (2%).
Preferred nesting trees
The Indochinese Roller’s nests
were opportunistically recorded in northeast India (n = 38) during 2005 to
2022. Fourteen nest attempts were observed at Buxa
Tiger Reserve, West Bengal, 16 nest attempts were recorded at Rani, Assam on
the same Sal tree, and six attempts were recorded at Kaziranga
National Park, Assam. The tree species preferred were Sal, Mynah, and Koroi. The average height of these preferred trees
was 23.7 m and the nest height was 9.2 m.
Discussion
The Indochinese Roller is known
to breed in April and May (Ali & Ripley 1987) and our observations exactly
match these reports in northern Bengal and Assam. We missed the actual
incubation period observation as we avoided checking the nest cavity during
this period which might have disturbed the nesting birds (Barve
et al. 2020).
The Indian Roller is known to
nest at ‘moderate height’ in the holes in tree trunks as well as in the walls.
In the current study, we recorded the nest height as 11.6 m while the mean
height of nests on the nine trees was observed to be 9.2 m, ranging between
4.5–14.6 m.
The Indian Roller is known to
pounce on frogs, plunge in water for fish, and hunt insects. We noticed and
photographed similar behaviour in the Indochinese
Roller during the study period. The C.
benghalensis is known to hunt snakes (Biddulph
1937; Ali & Ripley 1987; Vishnu & Ramesh 2021), our study records the
same behaviour in case of C. affinis
as well. Studies of pellets have suggested that the entomofauna consists major
chunk of the food of Indian Roller in south India (Sivakumaran
& Thiyagesan 2003). As the prey capture and
feeding observations were direct observations, the slightly smaller sized male
capturing smaller prey like insects with higher frequency and the slightly
heavier female focusing on larger vertebrate prey reminds the reversed sex
dimorphism and prey preference in raptors (Schantz & Nilsson 1981),
although elaborate data is required in case of C. affinis.
The Indian Roller plays an
important role in agricultural pest control and similar role is played by the
Indochinese Roller as evident from the diet. Though these resident species have
stable populations and are included in the ‘Least Concern’ category of the IUCN
Red List of Threatened Species, the migratory species of roller in India, the
European Roller Coracias garrulus, suffers a moderate decline (Tucker &
Heath 1994). Hence it is high time to collect the baseline data on the
Indochinese Roller which remains overlooked so far.
Table 1. Prey items captured by the
Indochinese Roller pair.
|
Prey items |
Average biomass of prey item in
g |
Number of preys captured by
female |
Biomass of prey captured by
female in g |
Number of preys captured by
male |
Biomass of prey captured by
male in g |
Total prey biomass in g |
Total number of prey items |
|
Insects |
0.1 |
35 |
3.5 |
51 |
5.1 |
8.6 |
86 |
|
Spiders |
0.1 |
1 |
0.1 |
0 |
0 |
0.1 |
1 |
|
Frogs |
10 |
2 |
20 |
0 |
0 |
20 |
2 |
|
Rats |
15 |
1 |
15 |
0 |
0 |
15 |
1 |
|
Reptiles |
20 |
3 |
60 |
1 |
20 |
80 |
4 |
|
Unknown |
1 |
2 |
2 |
4 |
4 |
6 |
6 |
|
Total |
|
44 |
100.6 |
56 |
29.1 |
129.7 |
100 |
For
figure & images - - click here for full PDF
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