Journal of Threatened Taxa |
www.threatenedtaxa.org | 26 August 2022 | 14(8): 21619–21641
ISSN 0974-7907 (Online) | ISSN 0974-7893
(Print)
https://doi.org/10.11609/jott.7613.14.8.21619-21641
#7613 | Received 09 August 2021 | Final
received 20 June 2022 | Finally accepted 04 August 2022
Checklist of
Carabidae (Coleoptera) in the Chinnar Wildlife Sanctuary, a dry forest in the
rain shadow region of the southern Western Ghats, India
M.C. Sruthi 1 & Thomas K. Sabu 2
1 Entomology Research
Unit, Post Graduate & Research Department of Zoology, St. Joseph’s College,
Devagiri, Kozhikode, Kerala 673008, India.
2 Department of
Zoology, University of Calicut, Tenhipalam, Kerala 673635, India,
1 sruthimangichalil@gmail.com,
2 sabukthomas@gmail.com (corresponding author)
Editor: Anonymity requested. Date of publication: 26 August
2022 (online & print)
Citation: Sruthi, M.C. &
T.K. Sabu (2022). Checklist of Carabidae (Coleoptera) in the
Chinnar Wildlife Sanctuary, a dry forest in the rain shadow region of the
southern Western Ghats, India. Journal of Threatened Taxa 14(8): 21619–21641. https://doi.org/10.11609/jott.7613.14.8.21619-21641
Copyright: © Sruthi & Sabu 2022. Creative Commons
Attribution 4.0 International License.
JoTT allows unrestricted use, reproduction, and distribution of this
article in any medium by providing adequate credit to the author(s) and the
source of publication.
Funding: Council for
Scientific and Industrial Research.
Competing interests: The authors declare no
competing interests.
Author details: M.C. Sruthi works as CSIR-SRF under the guidance of Dr. Sabu K.
Thomas. Her area of specialisation is the taxonomy of Carabidae. Sabu K. Thomas, is currently a professor
interested in the ecology & taxonomy of dung beetles & Carabidae and
home invading nuisance pest, Luprops tristis. He is currently engaged
with updating the taxonomy of Indian Carabidae. He has discovered 18 new
Carabidae species and 11 dung beetle species.
Author contributons: SKT and MCS reviewed
the earlier works and discussed the distribution patterns. MCS conducted the
field studies and prepared the images and the specimens.
Acknowledgements: Financial support
received from the Council for Scientific and Industrial Research by the first
author (CSIR, Govt. of India; Fileno: 08/453(0007)/2017-EMR-I) and the
infrastructure facilities from DST SERB funded major research project to second
author are gratefully acknowledged. We thank and acknowledge the Kerala Forest
and Wildlife Department for sample collection permissions; Beulah Garner,
curator, BMNH London, Thierry Deuve, Azadeh Taghavian Azari, Curator, MNHN
Paris, Aleksey A. Gusakov, MS Zoologist, curator of Coleoptera Zoological
Museum of the Moscow Lomonosov State University, for locating and sending
photographs; Akhil S.V., Jithmon V.A., Divya M., Ashly Kurian, Shigina K.,
Aswathi S.B., and Nijisha K. (St. Joseph’s College, Devagiri, Kozhikode) for
their logistic support.
Abstract: The first report on
the composition of carabids from a natural forest in peninsular India as well
as from a dry forest belt in the rain shadow region of the Western Ghats is
provided, with data on the subfamilies, tribes, genera, species, geographic
range, collection techniques, and the relevant literature details for all the
listed species. Fifty-four species belonging to 11 subfamilies and 31 genera
were recorded. Harpalinae, Lebiinae, and Scaritinae with 15, 14, and seven
species, respectively, are the species-rich subfamilies. The species list also
includes two first records from India, four first records from southern
India, and six species endemic to the Western Ghats and Sri Lanka biodiversity
hot spot.
Keywords: Carabids, Eastern
slope, endemism, first Indian record, ground beetles, peninsular India.
INTRODUCTION
The family Carabidae
(ground beetles) is composed of over 34,000 species distributed among 1,927
genera worldwide. Carabids occupy most land habitats on nearly all continents
(Lorenz 2005). These beetles are abundant in the field and attract attention
with their peculiar shape and coloration. Adults and larvae of most ground
beetle species are generalized predators of insects and other invertebrates;
however, many species are herbivores, omnivores or scavengers (Allen 1979).
Carabids are generally seen under stones, wood, moss, and bark (Andrewes 1929;
Thiele 1977), are sensitive to their environment, and are commonly used as
biological indicators (Rainio & Niemelä 2003; Koivula 2011). They are
useful in controlling the population build-up of soil-dwelling insects like
ants and termites (Kumar & Rajagopal 1990) as these beetles feed on the
immature stage of soil and litter-dwelling insects.
The Western Ghats
(WG), a chain of mountains of southwestern India, is one of the last remaining
stretches of the biodiverse tropical wet evergreen rainforests in peninsular
India and is a global biodiversity hotspot (Myers et al. 2000). The eastern
slope of the WG relies heavily on the north-east monsoon (October–December) for
precipitation, as opposed to the western scarps that receive almost 80% of
their rainfall between May and August due to the south-west monsoon (Anu et al.
2009). This variance in monsoon dependence is hypothesized to have led to
phenological differences amongst some congeneric populations from the eastern
and western slopes (Janani et al. 2017; Chaitanya et al. 2018).
Consequently, the faunal composition greatly varies between various segments of
the WG as revealed by the vertebrate group studies (Vijayakumar et al. 2014;
Deepak et al. 2016; Garg et al. 2017). Vertebrate groups have received a great
deal of attention in ecological studies conducted in the WG but the same is not
the case for most arthropod groups. Limited data exists on most coleopteran
families in general from the WG including ground beetles (Carabidae). Most
ground-beetles in the southern WG are found to live under upper layers of the
soil below stones, lower layers of litter and woody debris, and dry dung of
mega herbivores, and most are crepuscular and nocturnal. Available data on the
taxonomy of ground beetles is based on the species reported in the classical
work of Andrewes (1930), which is placed under two subfamilies: Harpalinae and
Carabinae, following the earlier classification of the family and in the recent
checklists of subfamilies, Lebiinae, Pterostichinae, Panagaeinae, and Dryptinae
(Shiju & Sabu 2019; Divya & Sabu 2020; Jithmon & Sabu 2021) do not
cover the entire family. There is no comprehensive data to understand the
Carabidae groups present in a natural ecosystem in the WG. In this work, we
list all the Carabidae species that have been recorded from a well-protected
wildlife sanctuary in the dry eastern slope of the southern WG to provide
baseline data about the composition of carabids in a natural habitat. This
checklist should greatly facilitate taxonomic and ecological studies by
complying with the current scientific knowledge. It will provide data on the
subfamilies, tribes, genera, species, geographic range, collection techniques,
and the relevant literature for all the listed species. Synonymies for each
species are followed by Lorenz (2005, 2021). Furthermore, the checklist could
be used in practical conservation programs for monitoring habitat changes in
dry forests.
MATERIAL AND METHODS
Study area
Chinnar Wildlife
Sanctuary is located in the rain shadow region of the WG (Figure 1). The
Sanctuary falls under the Anamudi Elephant Reserve and is situated 18 km north
of Marayur of Devikulam Taluk in Idukki district of Kerala, located between
10.25–10.35 N & 77.1–77.26 E, covering a total area of 90.44 km². The
dominant vegetation is dry deciduous forests followed by scrub jungle and
patches of riparian forests linearly spread out along the hill folds (Thomas et
al. 2018). Annual rainfall ranges 300–500 mm, the bulk of the rainfall is
received from north-east monsoon during October to December and the rainy
season lasts for about one month leading to a prolonged dry season and a short
rainy season (Management plan of Chinnar Wildlife Sanctuary 2012–13 to 2021–22;
Sabu & Nithya 2016).
Methods
The collections of
beetles were done using light traps, pitfall traps, and hand picking from the
thorny scrub jungle (Chinnar), dry deciduous forest (Alampetty), and riparian
forest (Kootar) during the dry season (January–September) and the rainy and
post rainy wet season (October–December) in 2019–2020. We followed the
classification pattern provided in Lorenz (2005) for subfamilies, tribes,
genera, and species. Species-level identification was done with the aid of
taxonomic keys in Andrewes (1929, 1935), Habu (1973), Balkenohl (2001), Kataev
(2012, 2018), Shiju et al. (2012), Kataev & Wrase (2016), Roux et al.
(2016), Sabu (2018), Shiju (2018), Akhil (2019), Akhil & Sabu (2019), Akhil
et al. (2019), Jithmon (2020), and by comparing with the holotypes and verified
specimens available in the insect depository of Zoological Survey of India,
Western Ghats Regional Centre (ZSI-WGRC) Kozhikode station. Images were taken
using Leica M 205C stereo zoom microscope fitted with Leica MC 170 HD digital
camera. Collected specimens are deposited at ZSI-WGRC. The checklist is grouped
by order, family, subfamily, tribe, genera, and species, each of which is
arranged alphabetically.
Abbreviations used
id. “Idem” (the same;
as just mentioned) | @—First report from India | #—First report from southern
India | *—Endemic to the Western Ghats | Ssp.—Subspecies.
World Zoogeographical
Regions
AUR—Australian Region
| IAR—Indo-Australian Region | ORR—Oriental Region | PAR—Palaearctic Region.
Geographical symbols
AF—Afghanistan;
AST—Australia; BGD—Bangladesh; BT—Bhutan; CBD—Cambodia; CHN—China;
EAI—East Indies; FUJ—Fujian; GUA—Guangdong; GUI—Guizhou; GUX—Guangxi;
HAI—Hainan; HKG—Hong Kong; HUB—Hubei; HUN—Hunan; IDS—Indonesia; IN—Iran;
JA—Japan; JIX—Jiangxi; LAO—Laos; MAC—Macao; MLS—Malaysia; MM—Myanmar; NC—North
Korea; NEC—New Caledonia; NP—Nepal; PA—Pakistan; PP—Philippines; SC—South
Korea; SCH—Sichuan; SEA—South East Asia; SHG—Shanghai; SM—Samoa; SRL—Sri Lanka;
TAI—Thailand; TD—Tajikistan; TM—Turkmenistan; TWN—Taiwan; UZ—Uzbekistan; VTN—Vietnam; YUN—Yunnan.
RESULTS
A total of 54 species of
ground beetles were examined. The checklist, distribution of the recorded
species are given below.
Order Coleoptera
Family Carabidae
Latreille 1802
Subfamily Anthiinae
Bonelli 1813
Tribe Helluonini Hope
1838
i. Genus Macrocheilus Hope
1838
Macrocheilus Hope 1838: 166.
= Acanthogenius Reiche 1843
= Macrochilus Agassiz 1847
= Macrocheilidius Jeannel 1949
1. Macrocheilus
bensoni Hope 1838
Macrocheilus bensoni Hope 1838: 166;
Andrewes 1930: 208; Lorenz 2005: 512; Shiju et al. 2012: 100; Löbl &
Löbl 2017: 577.
= Carabus trimaculatus Olivier 1790 (non
Villers, 1789)
= Helluo quadrimacultus Guérin-Méneville 1840
= Helluo tripustulatus Guérin-Méneville 1843
(non Dejean, 1825)
= Macrochilus
quadripustulatus Schmidt-Göbel 1846
= Macrochilus infuscatus Bates 1892a
= Macrochilus benarensis
Jedlička 1963
= Macrochilus
bimaculatus Jedlička 1965
= Macrochilus
quadrimaculatus (Guérin-Méneville 1840)
= Macrochilus
trimaculatus (G.A. Olivier 1790)
Specimens examined (n =
3): SJC-ZOO-CWSSMC001–003, Alampetty, 1 ex, Light trap, 25.ii.2020; 1 ex, hand
picking, 26.ii.2020; Kootar, 1 ex, pitfall trap, 26.x.2019.
Distribution: ORR - India (Assam (Andrewes 1930: 208),
Kerala: Kozhikode, Chinnar, Thamarassery (Shiju et al. 2012: 100)); SRL
(Andrewes 1930: 208); MM (Andrewes 1930: 208); LAO (Andrewes 1930: 208); VTN
(Andrewes 1930: 208); PAR - FUJ; GUA; GUI; GUX; HAI; JIX; YUN (Löbl
& Löbl 2017: 577); HKG (Andrewes 1930: 208); IAR
- PP (Andrewes 1930: 208); MLS (Andrewes 1930: 208).
*2. Macrocheilus
chinnarensis Akhil et al. 2019
Macrocheilus chinnarensis
Akhil et al. 2019: 28–33.
Distribution: ORR- India (Kerala: Chinnar
(Akhil et al. 2019: 28–33)).
ii. Genus Omphra Dejean
1825
Omphra Dejean 1825: 168, 283;
Reiche 1843: 330; Lacordaire 1854: 94; Chaudoir 1872a: 140; Sloane 1914: 570;
Andrewes 1930: 236; Csiki 1932: 1577; Jedlička 1963: 511; Lorenz 2005: 511;
Zhao et al. 2008: 372; Shiju & Sabu 2012: 2 ; Akhil & Sabu 2021:
11.
3. Omphra pilosa (Klug
1834)
Omphra pilosa (Klug) Reiche 1843: 330;
Erichson 1847: 141; Redtenbacher 1867: 5; Chaudoir 1872a: 141; Putzeys 1875a:
45; Andrewes 1921a: 163; id. 1923b: 460; id. 1927: 101; id. 1930: 237; Csiki
1932: 1578; Jedlička 1963: 512; Lorenz
2005: 511; Zhao et al. 2008: 371; Shiju & Sabu 2012: 8; Löbl & Löbl 2017: 578.
Helluo pilosus Klug 1834: 71
= Galerita attelaboides
Fabricius 1801
= Helluo pilosus Klug 1834
Specimens examined (n =
23): SJC-ZOO-CWSSMC004–026, Chinnar, 2 exs, pitfall, 25.ii.2020; Alampetty, 4
exs, pitfall trap, 26.x.2019; 3 exs, hand picking, 26.x.2019; 7 exs, pitfall
trap, 25.ii.2020; 4 exs, hand picking, 25.ii.2020; Kootar, 3 exs, pitfall trap,
26.x.2019.
Distribution: ORR - India (Kerala:
Arakulam, Chempery, Chinnar, Alampetty, Kuttiyadi, Kozhikode, Malappuram,
Thodupuzha, Mahe (Shiju & Sabu 2012: 8)); SRL (Andrewes 1930: 237); PAR -
India (Himachal Pradesh; Uttarakhand (Löbl & Löbl 2017: 578)); PA (Löbl & Löbl 2017: 578).
Subfamily Brachininae
Bonelli 1810
Tribe Brachinini Bonelli
1810
iii. Genus Styphlomerus
Chaudoir 1875
Styphlomerus Chaudoir 1875: 87, 88;
Erwin 1970: 39.
4. Styphlomerus
striatus Akhil & Sabu 2019
Styphlomerus striatus
Akhil & Sabu 2019: 468.
Specimens examined (n =
2): SJC-ZOO-CWSSMC027–028, Alampetty, 2 exs, light trap, 26.x.2019.
Distribution: ORR - India (Tamil Nadu:
Rajapalayam, Ettimadai; Kerala: Tholpetty (Akhil & Sabu 2019: 468))
Subfamily Dryptinae
Bonelli 1810
Tribe Dryptini Bonelli
1810
iv. Genus Drypta Latreille
1796
Drypta Latreille 1796: 75;
Fabricius 1801: 230; Latreille 1810: 117; Dejean 1825: 182; Schmidt- Göbel
1846: 22; Lacordaire 1854: 79; Andrewes 1924b: 51; id. 1930: 157; Lorenz 2005:
503; Jithmon & Sabu 2021: 18560.
5. Drypta lineola MacLeay
1825
Drypta lineola MacLeay 1825: 27;
Dejean 1825: 184; Redtenbater 4; Chaudoir 1877: 262; Bates 1883: 279; id. 1891:
336; id. 1892a: 383; Heyne-Tasch 13.t.2.f.25; Bouchard 1903: 173; Andrewes
1919a: 167; id. 1924c: 469; id. 1923e (1924): 460; id. 1924b: 52; id. 1930:
158; Lorenz 2005: 503; Jithmon & Sabu 2021: 18562.
= Desera lineola (W.S. MacLeay
1825)
Specimens examined (n =
1): SJC-ZOO-CWSSMC029, Alampetty, 1 ex, light trap, 26.x.2019.
Distribution: ORR - Throughout southeastern
Asia (Andrewes 1930: 158) India (Tamil Nadu: Rajapalayam, Kadayam (Jithmon
& Sabu 2021: 18560); Kerala: Padinjarathara (Jithmon & Sabu 2021:
18560)); MM (Andrewes 1930: 158); PAR - TWN; YUN (Andrewes 1930: 158; IAR
- IDS (Andrewes 1930: 158); PP (Andrewes 1930: 158); MLS (Andrewes 1930:
158).
Subfamily harpalinae Bonelli 1810
Tribe Anisodactylini
Lacordaire 1854
v. Genus Pseudognathaphanus
Schauberger 1932
Pseudognathaphanus Schauberger 1932: 57; Habu 1973: 62;
Noonan 1973: 344; id. 1976: 12; Löbl & Smetana 2003: 363; Lorenz 2005: 351;
Park et al. 2006: 96; Kataev & Wrase
2016: 224; Löbl & Löbl 2017: 508.
= Hiekea Ito 1997
= Protognathus Basilewsky 1950
6. Pseudognathaphanus
rusticus (Andrewes 1920)
Pseudognathaphanus
rusticus (Andrewes) Löbl & Smetana
2003: 363; Lorenz 2005: 351; Kataev & Wrase 2016: 232; Löbl & Löbl
2017: 508.
Gnathaphanus rusticus Andrewes 1920a: 107; id. 1924b:
30; id. 1930: 172; Kushwaha & Hegde 2015: 403.
= Gnathaphanus
rusticus Andrewes 1920
Specimens examined (n =
1): SJC-ZOO-CWSSMC030, Kootar, 1 ex, light trap, 26.ii.2020.
Distribution: ORR - India (New Delhi: Pusa;
Uttar Pradesh: Lucknow; Bihar: Chapra, Muzaffarpur, Purnea, Patna, Samastipur;
Madhya Pradesh; Odisha: Surada; Gujarat: Surat (Andrewes 1930: 172);
Maharashtra: Mumbai, Pune (Kataev & Wrase 2016: 232), Chikalda, Nagpur (Andrewes 1930: 172); Goa (Kataev & Wrase
2016: 232); Karnataka: Belgaum, Dharwar, North Karnataka (Andrewes 1930: 172); SRL (Andrewes 1930: 172); PAR -
India (Uttarakhand: Dehradun, Haridwar and Roorkee (Andrewes 1930: 172)), NP; PA
(Löbl & Löbl 2017: 508).
Tribe Stenolophini Kirby 1837
vi. Genus Stenolophus Dejean 1821
Stenolophus Dejean 1821: 15; id.
1829: 405; Lacordaire 1854: 303; Sloane 1898: 456; Tschitschérine 1900a:
364; id. 1901: 246; Andrewes 1924b: 40; id. 1930: 316; Habu 1973: 341; Noonan
1976: 17; Saha 1995: 67; Saha & Halder 2000: 15; Löbl &
Smetana 2003: 404; Lorenz 2005: 353; Park et al. 2006: 96; Löbl & Löbl
2017: 573.
# 7. Stenolophus
bajaurae Andrewes 1924
Stenolophus bajaurae Andrewes 1924b: 95; id.
1926a: 69; id. 1930: 316; Kataev 2002: 724; Löbl & Smetana 2003: 405;
Lorenz 2005: 354; Wrase 2005: 852; Kataev 2015: 93; id. 2015: 539; Kushwaha
& Hegde 2015: 401; Jaeger & Ahmed 2017: 613; Kataev 2002: 724;
Löbl & Löbl 2017: 574.
= Egadroma bajaurae (Andrewes
1924)
Specimens examined (n =
1): SJC-ZOO-CWSSMC031, Kootar, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Delhi (Kushwaha & Hegde
2015: 401); Uttar Pradesh: Fyzabad (Andrewes 1930: 316); Jharkhand:
Sarju valley (Andrewes 1930: 316)); PAR - India (Jammu-Kashmir (Andrewes 1930:
316), Himachal Pradesh: Kangra, Bajaura, Spiti, Manikaran
(Andrewes 1930: 316); Uttarakhand: Kumaon (Andrewes 1930:
316)); AF; NP; PA; TD; TM; UZ (Löbl & Löbl 2017: 574).
@8. Stenolophus
lucidus Dejean 1829
Stenolophus lucidus Dejean 1829: 419;
Andrewes 1930: 317; Löbl & Smetana 2003: 405; Lorenz 2005: 355; Löbl
& Löbl 2017: 574.
= Egadroma lucida (Dejean
1829)
Specimens examined (n =
1): SJC-ZOO-CWSSMC032, Kootar, 1 ex, light trap, 26.ii.2020.
Distribution: ORR - EAI (Andrewes 1930:
317); PAR - BT; FUJ; GUA; GUX; HAI; TWN; YUN;
JA; NP (Löbl & Löbl 2017: 574).
9. Stenolophus
quinquepustulatus (Wiedemann 1823)
Stenolophus quinquepustulatus
(Wiedemann) Dejean 1829: 414; Bates
1873: 270; Putzeys 1875a: 49; Bates
1889: 272; id. 1891: 333; Bouchard 1903: 172; Lesne 1904: 76; Sloane 1920a:
321; Andrewes 1921a: 171; id. 1924c: 469; id. 1930: 317; Habu 1973: 382; Saha 1995: 68; Löbl
& Smetana 2003: 405; Lorenz 2005: 355; Park et al. 2006: 96; Jaeger &
Ahmed 2017: 614; Löbl & Löbl 2017: 574.
= Badister
quinquepustulatus Wiedemann 1823
= Stenolophus
rectifrons Bouchard 1903 (non Bates 1892)
= Stenolophus
connexus Schauberger 1928
= Stenolophus
apicalis Jedlička 1952
= Stenolophus
tripustulatus Jedlička 1952
= Stenolophus
conjunctus Jedlička 1956
= Stenolophus
unipustulatus Jedlička 1952
= Acupalpus connexus (Schauberger 1928)
= Egadroma
quinquepustulata (Wiedemann 1823)
Specimens examined (n =
2): SJC-ZOO-CWSSMC033–34, Kootar, 2 exs, Light trap, 26.ii.2020.
Distribution: ORR - India (Uttar Pradesh; West
Bengal: Singur, Hooghly (Saha 1995: 68)); MM (Habu 1973: 382); SRL (Habu 1973: 382);
TAI (Habu 1973: 382); VTN (Park et al. 2006: 96); PAR - FUJ; GUI; GUX; HAI;
HKG; HUB; HUN; JIX; MAC; TWN; YUN; NP; SC; SCH; SHG (Löbl & Löbl 2017: 574)); JA (Habu 1973: 382); PA (Habu 1973: 382); IAR - SM (Habu 1973: 382); IDS (Habu 1973: 382); MLS (Habu 1973: 382); PP (Habu 1973: 382); AUR - AST (Habu 1973: 382).
10. Stenolophus
smaragdulus (Fabricius 1798)
Stenolophus smaragdulus (Fabricius) Bates 1886:
80; id. 1891: 333; id. 1892a: 349; Bouchard 1903: 172; Sloane 1920a: 321;
Andrewes 1921a: 160; id. 1924b: 40; id. 1930: 318; Habu 1973: 377;
Saha 1995: 69; Saha & Halder 2000: 16; Löbl & Smetana 2003: 405; Lorenz
2005: 355; Park et al. 2006: 96; Jaeger & Ahmed 2017: 614; Löbl & Löbl
2017: 575.
Carabus smaragdulus Fabricius 1798: 60; id.
1801: 209; Dejean 1829: 418; Hope 1838: 93; Schaum 1847: 49; Motschulsky 1855:
43.
= Carabus smaragdulus Fabricius 1798
= Egadroma smaragdula Motschulsky 1864
= Harpalus trechoides Hope 1845
= Harpalus stolidus Walker 1858
= Egadroma apicalis Motschulsky 1864
= Stenolophus
transmutans Bates 1886
= Stenolophus chalceus Lesne 1904 (non Bates
1873)
= Egadroma smaragdula (Fabricius 1798)
= Stenolophus apicalis (Motschulsky 1864)
= Stenolophus
stolidus (Walker 1858)
= Stenolophus
trechoides (Hope 1845)
Specimens examined (n =
1): SJC-ZOO-CWSSMC035, Kootar, 1 ex, light trap, 25.ii.2020.
Distribution: Throughout the whole of
Southeast Asia extending from JA in the North to Queensland in South
(Andrewes 1930: 318); ORR - India (West Bengal: Kolkata, Kharagpur,
Purulia, Medinipur (Saha 1995: 69); Meghalaya: Khasi, Jayantia Hill (Saha &
Halder 2000: 16)); MM (Habu 1973: 377); SRL (Habu 1973: 377); TAI (Habu 1973: 377); VTN (Park et al. 2006: 96); PAR - India (Himachal Pradesh (Löbl & Löbl 2017:
575); West Bengal: Darjeeling District (Saha 1995: 69));
BT; FUJ; GUA; HAI; HKG; JIX; MAC; NP; PA; TWN; YUN (Löbl & Löbl 2017: 575);
JA (Habu 1973: 377); IAR - IDS (Habu 1973: 377); MLS (Habu 1973:
377); PP (Habu 1973: 377); AUR - AST
(Habu 1973: 377).
Tribe Harpalini Bonelli
1810
vii. Genus Allosiopelus
Ito 1995
Allosiopelus Ito 1995: 153; Lorenz 2005: 376.
11. Allosiopelus
punctatipennis Ito 1995
Allosiopelus
punctatipennis Ito 1995: 154; Lorenz 2005: 376.
Specimens examined (n =
2): SJC-ZOO-CWSSMC036–037, Alampetty, 2 exs, light trap, 26.x.2019.
Distribution: ORR - India (Tamil Nadu: Tharangambadi; Pondicherry (Ito
1995: 154)).
viii. Genus Amblystomus
Erichson 1837
Amblystomus Erichson 1837: 59;
Lacordaire 1854: 301; Reitter 1883: 139; Tschitschérine 1900a: 348; Sloane
1920b: 131; Andrewes 1924b: 33; id. 1930: 17; Habu 1973: 15; Noonan 1976: 54; Saha
1995: 56; Löbl & Smetana 2003: 360; Lorenz 2005: 384; Park
et al. 2006: 95; Löbl & Löbl 2017: 502.
= Hispalis Rambur 1838
= Artizoum Gistel 1857
= Megaristerus Nietner 1858
= Notophilus Blackburn 1888
= Thenarotidius Sloane 1898
= Psilonothus Sloane 1900
= Entomorrhinus Jeannel 1948
@ 12. Amblystomus
aenescens (Motschulsky 1858)
Amblystomus aenescens (Motschulsky)
Andrewes 1928: 21; id. 1930: 17; id. 1933: 7; Lorenz 2005: 384.
= Hispalis aenescence Motschulsky 1858
Specimens examined (n =
4): SJC-ZOO-CWSSMC038–041, Alampetty, 3 exs, light trap, 26.ii.2020; 1 ex,
pitfall trap, 26.ii.2020.
Distribution: ORR - EAI (Andrewes 1930:
17).
13. Amblystomus
fuscescens (Motschulsky 1858)
Amblystomus
fuscescens (Motschulsky) Bates 1892a: 334; Lesne 1904: 73;
Andrewes 1919a: 198; id. 1928: 21; id. 1930: 18; Kapur 1945: 326; Lorenz
2005: 384.
= Hispalis fuscescens Motschulsky 1858
Specimens examined (n =
20): SJC-ZOO-CWSSMC042–061, Alampetty, 10 exs, light trap, 26.x.2019; 3 exs,
pitfall trap, 26.x.2019; 2 exs, hand picking, 26.x.2019; 3 exs, light trap,
25.ii.2020; 1 ex, pitfall trap, 25.ii.2020; 1 ex, hand picking, 25.ii.2020.
Distribution: ORR - India (Assam;
Manipur: Imphal Valley; Karnataka: Mysore (Kapur 1945: 326)); EAI (Andrewes 1930: 18); SRL (Andrewes 1930: 18);
MM (Andrewes 1930: 18); TAI (Andrewes 1930: 18).
# 14. Amblystomus
indicus (Nietner 1858)
Amblystomus indicus (Nietner) Bates 1886:
76; id. 1889: 271; id. 1891: 331; id.
1892a: 336; id. 1892b: 231; Sloane 1920a: 321; Andrewes 1927: 103; id. 1930:
19; Lorenz 2005: 384; Kushwaha & Hegde 2015: 402; Löbl
& Löbl 2017: 502.
= Megaristerus indicus Nietner 1858
= Entomorrhinus
indicus (Nietner 1858)
Specimens examined (n =
19): SJC-ZOO-CWSSMC062–80, Alampetty, 7 exs, light trap, 26.x.2019; 2 exs,
pitfall trap, 26.x.2019; 3 exs, hand picking, 26.x.2019; 6 exs, light trap,
25.ii.2020; 1 ex, hand picking, 25.ii.2020.
Distribution: ORR - India (Uttar
Pradesh: Jalaun, Orai, Jhansi; Madhya Pradesh: Pathrora (Kushwaha & Hegde
2015: 402); Jharkhand: Chota Nagpur, Tetara (Andrewes 1930:
19)); MM (Kushwaha & Hegde 2015: 402); VTN (Kushwaha & Hegde
2015: 402); SRL (Andrewes 1930: 19); AUR - AST (Andrewes
1930: 19).
ix. Genus Dioryche
MacLeay 1825
Dioryche MacLeay 1825: 21; Lacordaire
1854: 300; Bates 1873: 271; Alluaud 1917: 321; Andrewes 1919a: 156; id.
1924b: 32; id. 1930: 146; Noonan 1976: 47; id. 1985: 34; Saha
1995: 62; Löbl & Smetana 2003: 369; Lorenz 2005: 376; Kataev 2012: 112; Kushwaha & Hegde
2015: 402; Löbl & Löbl 2017: 518.
= Hypodioryche
Schauberger 1935
15. Dioryche
cuprina (Dejean 1829)
Dioryche cuprina (Dejean) Kataev 2012: 114; Löbl
& Löbl 2017: 518.
= Selenophorus cuprinus Dejean 1829
= Harpalus
colombensis Nietner 1857a
= Cardiaderus scitus Walker 1858
= Dioryche
colombensis (Nietner 1857)
= Dioryche scita (Walker
1858)
= Selenophorus
colombensis (Nietner 1857)
Specimens examined (n =
2): SJC-ZOO-CWSSMC081–082, Alampetty, 2 exs, light trap, 26.x.2019.
Distribution: ORR - India (Goa ;
Karnataka : Kanara ; Tamil Nadu: Chennai, Kariakal, Coimbatore; Pondicherry; Kerala:
Thiruvananthapuram, Mahe, Kozhikode, Kallar (Kataev 2012: 114)); SRL (Kataev 2012: 114); TAI (Kataev 2012: 114); PAR - NP (Kataev 2012: 114); PA (Löbl & Löbl 2017: 518).
16. Dioryche
dravidana Kataev 2012
Dioryche dravidana Kataev 2012: 123.
Specimens examined (n =
1): SJC-ZOO-CWSSMC083, Alampetty, 1 ex, pitfall trap, 26.x.2019.
Distribution: ORR - India (Karnataka:
Mysore, Shimoga; Tamil Nadu: Shambaganur, Madura (Kataev 2012: 123)).
17. Dioryche torta
MacLeay 1825
Dioryche torta MacLeay 1825: 21;
Hope 1838: T. 2; Bates 1873: 271; Andrewes 1919a: 154; id. 1926a: 68; id. 1930: 148; Noonan 1985: 35; Saha
1995: 63; Lorenz 2005: 376; Löbl & Smetana 2003: 369; Lorenz
2005: 376; Löbl & Löbl 2017: 518.
Specimens examined (n =
2): SJC-ZOO-CWSSMC084–085, Alampetty, 1 ex, pitfall trap, 26.x.2019; 1 ex,
light trap, 25.ii.2020.
Distribution: ORR - All the Indian States
(Saha 1995: 63) India (West Bengal: Murshidabad (Saha 1995: 63));
SRL (Andrewes 1930: 148); MM (Andrewes 1930: 148); PAR - GUA; HAI; NP; PA;
YUN (Löbl & Löbl 2017: 518); IAR - IDS (Andrewes 1930: 148).
x. Genus Ophoniscus
Bates 1892
Ophoniscus Bates 1892a: 337;
Andrewes 1923b: 446; id. 1930: 242; id. 1939: 136; Noonan 1976: 46;
id. 1985: 31; Saha 1995: 63; Löbl & Smetana 2003: 388; Kataev 2005: 269;
Lorenz 2005: 376; Kataev & Wrase 2012: 215; Löbl & Löbl 2017: 546;
Kataev 2018: 319.
*18. Ophoniscus
puneensis Kataev 2018
Ophoniscus puneensis Kataev 2018: 321.
Specimens examined (n =
1): SJC-ZOO-CWSSMC086, Alampetty, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Maharashtra:
Mulshi environment (Kataev 2018: 321)).
xi. Genus Parophonus
Ganglbauer 1891
Parophonus Ganglbauer 1891a: 340; Jeannel 1942: 625;
Noonan 1976: 45; id. 1985: 19; Löbl & Smetana 2003: 392; Lorenz 2005: 373;
Kataev 2010: 278; Löbl & Löbl 2017: 553.
19. Parophonus
acutangulus (Bates 1891)
Parophonus acutangulus (Bates) Andrewes 1930: 184;
Kataev 2010: 296; Löbl & Löbl 2017: 553.
= Hypolithus
acutangulus Bates 1891
= Hyperpalus gracilis Andrewes 1947
= Parophonus gracilis (Andrewes
1947)
= Trichotichnus
javanus (Gory 1833)
Specimens examined (n =
1): SJC-ZOO-CWSSMC087, Alampetty, 1 ex, light trap, 26.x.2019.
Distribution: ORR - India (Delhi;
Uttar Pradesh: Allahabad, Sitapur; Jharkhand: Chota
Nagpur- Tetara; Madhya Pradesh: Mhow; Gujarat: Surat; Maharashtra: Mumbai; Tamil Nadu:
Coimbatore, Tharangambadi (Andrewes 1930: 184)); MM (Kataev 2010: 296); SRL (Andrewes 1930: 184); PAR - India
(Jammu Kashmir (Kataev 2010: 296); Uttarakhand: Dehra Dun
(Andrewes 1930: 184); West Bengal: Barodabri (Kataev 2010: 296)); NP (Kataev 2010: 296); PA (Kataev 2010: 296); IAR - IDS (Andrewes 1930: 184).
20. Parophonus
indicus (Andrewes 1931)
Parophonus indicus (Andrewes) Noonan 1985: 22;
Lorenz 2005: 374; Kataev 2010: 283 ; Löbl & Löbl 2017: 553.
= Hyparpalus indicus Andrewes 1931a
= Hypolithus
cyaneotinctus Bates 1891 [non Bates 1889]
= Trichotichnus
indicus (Andrewes 1931)
Specimens examined (n =
1): SJC-ZOO-CWSSMC088, Alampetty, 1 ex, light trap, 26.x.2019.
Distribution: ORR - India (Uttar Pradesh; Bihar: Monghyr;
Jharkhand: Chota Nagpur-Tetara, Barwa, Konbir, Ranchi; Madhya Pradesh:
Balaghat, South Mandla (Andrewes 1931a: 516), Motinala, Seoni, Khawasa (Kataev
2010: 283); Karnataka: Mysore, Bangalore, Nandidrug, Chikkaballapura (Andrewes
1931a: 516)); SRL (Kataev 2010: 283); PAR - India (Jammu Kashmir (Kataev 2010: 283); Uttarakhand: Dehra Dun
(Andrewes 1931a: 516); Sikkim (Andrewes 1931a: 516)); PA (Kataev 2010: 283).
Subfamily Lebiinae
Bonelli 1810
Tribe Cyclosomini
Laporte De Castelnau 1834
xii. Genus Cyclicus Jeannel
1949
Cyclicus Jeannel 1949: 865, 870;
Basilewsky 1953: 117; id. 1956: 464; Lorenz 2005: 452.
= Metacyclicus Jeannel 1949
21. Cyclicus elegans (Andrewes
1931)
Cyclicus elegans (Andrewes) Lorenz 2005:
452; Shiju & Sabu 2019: 11.
= Tetragonoderus elegans
Andrewes 1931a
Specimens examined (n =
13): SJC-ZOO-CWSSMC089–101, Chinnar, 2 exs, light trap, 26.x.2019; Kootar, 3
exs, light trap, 26.x.2019; 4 exs, pitfall trap, 26.x.2019; 2 exs, hand picking,
26.x.2019; 2 exs, pitfall trap, 25.ii.2020.
Distribution: ORR - India (Kerala:
Charalmedu, Nedumkayam (Shiju & Sabu 2019: 11)); PAR - India
(Uttarakhand: Bindal River, Chakata Range, Dehra Dun, Deoba Nadi River,
Hathibarkala, Kali Valley, Nandhaur River, West Almora (Andrewes 1931a: 524)).
22. Cyclicus
fimbriatus (Bates 1886)
Cyclicus fimbriatus (Bates) Lorenz 2005:
452; Shiju & Sabu 2019: 11.
Tetragonoderus
fimbriatus Bates 1886: 202; Andrewes 1930: 344; Löbl & Löbl
2017: 498.
= Tetragonoderus punctatus
Schmidt-Göbel 1846 (non Wiedemann 1823)
= Cyclicus fimbriatus (Bates
1886)
Specimens examined (n =
1): SJC-ZOO-CWSSMC102, Alampetty, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Karnataka:
North Karnataka, Belgaum, Managanali, Mysore- Teppukadu
(Andrewes 1930: 344); Tamil Nadu: Nilgiri Hills-Hill Grove (Andrewes 1930:
344), Srivilliputhur (Shiju & Sabu 2019: 11), Tiruchirappally
(Andrewes 1930: 344); Kerala: Bhawani Valley (Andrewes 1930: 344), Kozhikode,
Nedumkayam (Shiju & Sabu 2019: 11)); SRL (Andrewes
1930: 344); MM (Andrewes 1930: 344); PAR - CHN (Löbl & Löbl 2017:
498).
xiii. Genus Tetragonoderus
Dejean 1829
Tetragonoderus Dejean 1829: 485;
Schmidt-Göbel 1846: 92; Lacordaire 1854: 132; Chaudoir 1876a: 33; Horn 1882:
127; Andrewes 1924b: 60; id. 1930: 343; Blackwelder 1944: 52; Jeannel 1949:
865; Basilewsky 1956: 463; Jedlička 1963: 291; Saha et al. 1992: 49; Lorenz
2005: 453; Löbl & Löbl 2017: 498.
23. Tetragonoderus
notaphioides Motschulsky 1861
Tetragonoderus notaphioides
Motschulsky 1861: 99; Chaudoir 1876a: 54; Bates 1886: 201; Andrewes 1928: 24;
id. 1930: 345; Lorenz 2005: 453; Shiju & Sabu 2019: 12.
Specimens examined (n =
2): SJC-ZOO-CWSSMC103–104, Kootar, 2 exs, pitfall trap, 26.x.2019.
Distribution: ORR - India (Odisha: Berhampur,
Puri, Rambha- Ganjam, Barkuda Island- Chilka Lake; Maharashtra: Bhandara,
Karnataka: North Karnataka; Tamil Nadu: Chennai, Tiruchirappally, Thrangambadi,
Palni Hills (Andrewes 1930: 345); Kerala: Kozhikode, Ambalavayal (Shiju &
Sabu 2019: 12)); SRL (Andrewes 1930: 345).
Tribe Lebiini Bonelli
1810
xiv. Genus Anchista
Nietner 1857
Anchista Nietner 1857c: 523; id.
1857b: 374; Chaudoir 1877: 236; Andrewes 1926b: 346; id. 1930: 22; Csiki 1932:
1455; Jedlička 1963: 449; Habu 1967: 137; Darlington 1968: 139; id. 1970: 45;
Habu 1982: 102; Kirschenhofer 1994: 1006; Lorenz 2005: 491; Löbl & Löbl
2017: 623.
= Paraphaea Bates 1873
24. Anchista
fenestrata (Schimdt-Göbel 1846)
Anchista fenestrata (Schmidt-Göbel)
Chaudoir 1872a: 168; Bates 1892a: 424; Andrewes 1923a: 20; id. 1930: 23; Csiki
1932: 1456; Jedlička 1963: 449; Lorenz 2005: 491; Shi et al. 2013: 27; Löbl
& Löbl 2017: 623; Shiju & Sabu 2019: 40.
= Plochionus
fenestrata Schmidt-Göbel 1846
Specimens examined (n =
15): SJC-ZOO-CWSSMC105–119, Chinnar, 1 ex, light trap, 26.x.2019; Alampetty, 6
exs, light trap, 26.x.2019; 4 exs, light trap, 25.ii.2020; Kootar, 3 exs,
26.x.2019; 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Rajasthan;
Bihar; Jharkhand: Singbhum (Andrewes 1930: 23); Karnataka: Gundelpet (Shiju
& Sabu 2019: 40); Tamil Nadu: Alwarkurichi, Srivilliputhur, Thambaram
(Shiju & Sabu 2019: 40); Pondicherry (Andrewes 1930: 23); Kerala:
Charalmedu, Chinnar-Alampetty; Koorachundu, Nedumkayam, Thamarassery (Shiju
& Sabu 2019: 40)); SRL (Andrewes 1930: 23); MM (Andrewes 1930: 23); PAR
- India (Uttarakhand: Dehra Dun; West Bengal); NP (Löbl & Löbl 2017:
623).
xv. Genus
Anomotarus Chaudoir 1875
Anomotarus Chaudoir 1875: 48;
Sloane 1917: 435; id. 1920b: 170; Andrewes 1930: 27; Jedlička 1963: 450; Lorenz
2005: 497; Löbl & Löbl 2017: 580.
25. Anomotarus
stigmula (Chaudoir 1852)
Anomotarus stigmula (Chaudoir) Andrewes
1930: 28; Jedlička 1963: 451; Lorenz 2005: 497; Löbl & Löbl 2017: 580;
Shiju & Sabu 2019: 42.
= Cymindis stigmula Chaudoir 1852
Specimens examined (n =
1): SJC-ZOO-CWSSMC120, Alampetty, 1 ex, light trap, 26.x.2019.
Distribution: ORR - India (Assam:
Gauhati (Andrewes 1930: 28); Maharashtra: Mumbai- Khandesh, Nagpur; Karnataka:
Belgaum (Andrewes 1930: 28), Gundelpet (Shiju & Sabu 2019: 42), Mysore-
Nandidurg; Tamil Nadu: Chennai (Andrewes 1930: 28), Srivilliputhur (Shiju &
Sabu 2019: 42); Kerala: Charalmedu, Eravikulam National Park, Koorachundu,
Nedumkayam, Thamarassery, Vazhachal, Vettiozhinjathottam (Shiju & Sabu
2019: 42)); MM (Andrewes 1930: 28); SRL (Andrewes 1930: 28); PAR - India
(Himachal Pradesh (Löbl & Löbl 2017: 580); Uttarakhand: Dehra Dun (Andrewes
1930: 28)); JA (Andrewes 1930: 28); NP; PA (Löbl & Löbl 2017: 580); TWN
(Jedlička 1963: 451); IAR - IDS (Andrewes 1930: 28); NEC (Andrewes 1930:
28).
xvi. Genus Apristus
Chaudoir 1846
Apristus Chaudoir 1846: 62;
Lacordaire 1854: 123; Horn 1882: 133; Andrewes 1930: 33; Ganglbauer 1892: 397
& 401; Jedlička 1933a: 87; Blackwelder 1944: 59; Jedlička 1963: 427;
Gueorguiev & Gueorguiev 1995: 32 & 229; Kryzhanovskij et al. 1995: 165;
Lorenz 2005: 472; Park et al. 2006: 100; Löbl & Löbl 2017: 595.
= Crepnos
Baudi Di Selve 1864
= Crephnos
Jakobson 1908
26. Apristus
aeneipennis (Schmidt-Göbel 1846)
Apristus aeneipennis (Schmidt-Göbel)
Chaudoir 1850: 67; Motschulsky 1855: 50; Fairmaire 1888: 335; Andrewes 1923a:
15; id. 1930: 33; Jedlička 1963: 430; Lorenz 2005: 472; Park et al. 2006: 100;
Shiju & Sabu 2019: 26.
= Lionychus
aeneipennis Schmidt-Göbel 1846
Specimens examined (n =
1): SJC-ZOO-CWSSMC121, Alampetty, 1 ex, hand picking, 26.x.2019.
Distribution: ORR - India (Maharashtra:
Lonavla; Karnataka: Mysore-Teppukadu (Andrewes 1930: 33)); MM (Andrewes 1930:
33); VTN (Andrewes 1930: 33).
27. Apristus
subtransparens Motschulsky 1861
Apristus subtransparens Motschulsky 1861: 104;
Bates 1886: 206; id. 1892b: 233; Andrewes 1928: 21; id. 1930: 34; Lorenz 2005:
472; Löbl & Löbl 2017: 596; Shiju & Sabu 2019; 27.
Specimens examined (n =
2): SJC-ZOO-CWSSMC122–123, Kootar, 2 exs, hand picking, 26.x.2019.
Distribution: ORR - India (Kerala: Chinnar,
Koottar, Nedumkayam, Thamarassery (Shiju & Sabu 2019; 27)); SRL (Andrewes
1930: 34); NP; PA (Löbl & Löbl 2017: 596).
xvii. Genus
Catascopus Kirby 1825
Catascopus Kirby 1825: 94;
Latreille et Dejean 1824: 115; Macleay 1825: 14; Dejean 1825: 328;
Schmidt-Göbel 1846: 80; Lacordaire 1854: 145; Chaudoir 1861: 116; id. 1872b:
244; Andrewes 1924b: 62; id. 1926b: 348; id. 1930: 74; id. 1931b: 62; id. 1937:
187; Jedlička 1935: 9; Jeannel 1942: 1017; Blackwelder 1944: 57; Basilewsky
1956: 485; Jedlička 1963: 379; Lorenz 2005: 454; Löbl & Löbl 2017: 620.
28. Catascopus
cingalensis Bates 1886
Catascopus cingalensis
Bates 1886: 203; Andrewes 1924b: 117; id. 1930: 75; Lorenz 2005: 454; Shiju
& Sabu 2019: 15.
= Catascopus reductus Chaudoir 1861
= Catascopus severini Bates 1891
Specimens examined (n =
1): SJC-ZOO-CWSSMC124, Chinnar, 1 ex, hand picking, 26.x.2019.
Distribution: ORR - India (Jharkhand: Chota
Nagpur- Tetara; Madhya Pradesh: Mhow; Odisha: Surada; Karnataka: Chikkaballapura;
Tamil Nadu: Nilgiri Hills (Andrewes 1930: 75)); SRL (Andrewes 1930: 75).
29. Catascopus cyanellus
Chaudoir 1848
Catascopus cyanellus Chaudoir 1848: 113; id.
1861: 118; Andrewes 1930: 75; Lorenz 2005: 454; Löbl & Löbl 2017: 620;
Shiju & Sabu 2019 : 15.
= Catascopus reductus Walker 1858
Specimens examined (n =
7): SJC-ZOO-CWSSMC125–131, Chinnar, 2 exs, pitfall trap, 26.x.2019; 5 exs, hand
picking, 26.x.2019.
Distribution: ORR - India (Maharashtra:
Dapoli; Karnataka : North Karnataka; Tamil Nadu: Coimbatore (Andrewes
1930: 75)); PAR - India (Uttarakhand: Dehra Dun (Andrewes 1930: 75)); NP
(Andrewes 1930: 75).
xviii. Genus Lebia Latreille
1802
Lebia Latreille 1802: 85;
Dejean 1825: 253; Schmidt-Göbel 1846: 43; Lacordaire 1854: 127; Chaudoir 1871a:
111–255; id. 1871b: 1–87; Horn, 1882: 130; Fowler 1887: 136; Ganglbauer 1892:
397; Silvestri 1904: 68–84; Andrewes 1930: 191; Alluaud 1936: 8; Jedlička
1933b: 144; Jeannel 1942: 1028; id. 1949: 882, 902; Jedlička 1963: 314;
Blackwelder 1944: 52; Mateu 1984: 398; Gueorguiev & Gueorguiev 1995: 31,
221; Kryzhanovskij et al. 1995: 161; Hůrka 1996: 468, 470; Lorenz 2005: 481;
Park et al. 2006: 102; Löbl & Löbl 2017: 611.
30. Lebia baconi (Chaudoir
1871)
Lebia baconi
(Chaudoir) Andrewes 1930: 191; Lorenz 2005: 487; Löbl & Löbl 2017: 616;
Shiju & Sabu 2019 : 37.
= Nematopeza baconi Chaudoir 1871a
Specimens examined (n =
1): SJC-ZOO-CWSSMC132, Alampetty, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Bihar: Chapra;
Madhya Pradesh: Hoshangabad (Andrewes 1930: 191); Tamil Nadu: Srivilliputhur
(Shiju & Sabu 2019: 37)).
31. Lebia calycophora
Schmidt-Göbel 1846
Lebia (Poecilothais)
calycophora Schmidt-Göbel 1846: 44; Bates 1892a: 427; Andrewes
1923a: 21; id. 1930: 191; Jedlička 1963: 322–325; Lorenz 2005: 488; Park et al.
2006: 102; Löbl & Löbl 2017: 616; Shiju & Sabu 2019: 37.
= Lebia comitata Bates 1873
= Lebia farai Jedlička 1951
Specimens examined (n =
3): SJC-ZOO-CWSSMC133–135, Alampetty, 2 exs, light trap, 26.x.2019; Kootar, 1
ex, light trap, 25.ii.2020.
Distribution: ORR - India (Nagaland: Naga
Hills; Assam: Khasi Hills, Patkai Hills (Andrewes 1930: 191); Kerala: Aralam
(Shiju & Sabu 2019: 37)); MM (Andrewes 1930: 191); TAI (Andrewes 1930:
191); VTN (Jedlička 1963: 322–325); PAR - CHN (Jedlička 1963: 322–325);
FUJ; HUN; PA; TWN (Löbl & Löbl 2017: 616); IAR - IDS (Jedlička 1963:
322–325); MLS (Jedlička 1963: 322–325).
32. Lebia indica
Liebke 1938
Lebia indica Liebke 1938: 109;
Lorenz 2005: 487; Löbl & Löbl 2017: 616; Shiju & Sabu 2019: 37.
= Nematopeza decora
Chaudoir 1871c
= Lebia decora (Chaudoir 1871)
= Nematopeza indica (Liebke 1938)
Specimens examined (n =
1): SJC-ZOO-CWSSMC136, Alampetty, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Tamil Nadu:
Alwarkurichi, Sankarankovil (Shiju & Sabu 2019: 37))
Tribe Odacanthini
Laporte De Castelnau 1834
xix. Genus
Pentagonica Schmidt-Göbel 1846
Pentagonica Schmidt-Göbel 1846:
47; Lacordaire 1854: 133; Schaum 1863: 74; Bates 1873: 321; Chaudoir 1877: 212;
Sloane 1898: 494 & 513; Dupuis 1913a: 2; Andrewes 1926b: 353; id. 1930:
259; Jeannel 1942: 1017; Blackwelder 1944: 63; Jeannel 1949: 768; Basilewsky
1956: 472; Jedlička 1963: 505; Darlington 1968: 192; id. 1970: 46; Lorenz 2005:
445; Park et al. 2006: 103; Löbl & Löbl 2017: 640.
= Rhombodera Reiche 1842
= Didetus LeConte 1853
= Elliotia Nietner 1856
= Trichothorax Montrouzier 1860
= Xenothorax Wollaston 1867
= Wakefieldia Broun 1880
33. Pentagonica
ruficollis Schmidt-Göbel 1846
Pentagonica
ruficollis Schmidt-Göbel 1846: 48; Bates 1892a: 426; Dupuis
1913a: t. 5, f. 9⎼11; Andrewes 1923a:
23; id. 1926b: 353; id. 1930: 261; Jedlička 1963: 509; Lorenz 2005: 446; Park
et al. 2006: 104; Löbl & Löbl 2017: 641; Shiju & Sabu 2019: 8.
= Pentagonica dichroa Sloane 1903
Specimens examined (n =
2): SJC-ZOO-CWSSMC137–138, Alampetty, 1 ex, light trap, 26.x.2019; Chinnar, 1
ex, light trap, 25.ii.2020.
Distribution: ORR - India (Assam: Patkai
Hills; Tamil Nadu: Aratapara, Nilgiri Hills (Andrewes 1930: 261)); SRL
(Andrewes 1930: 261), MM (Andrewes 1930: 261); VTN (Andrewes 1930: 261); PAR
- GUA; HKG; YUN; NP; TWN (Löbl & Löbl 2017: 641); IAR - IDS
(Andrewes 1930: 261); AUR - AST (Andrewes 1930: 261).
34. Pentagonica venusta Andrewes 1933
Pentagonica venusta Andrewes 1933: 17;
Lorenz 2005: 446; Shiju & Sabu 2019: 8.
Specimens examined (n =
1): SJC-ZOO-CWSSMC139, Alampetty, 1 ex, light trap, 26.x.2019.
Distribution: ORR - India (Karnataka:
Belgaum, Coorg, Mysore- Nandidurg, South Mangalore; Tamil Nadu: Nilgiri
Hills-Kallar (Andrewes 1933: 17)); SRL (Andrewes 1933: 17).
Subfamily Licininae
Bonelli 1810
Tribe Chlaenini
Brulle 1834
xx. Genus Chlaenius
Bonelli 1810
Chlaenius MacLeay 1825: 13;
Dejean 1826: 297, 368; Schmidt-Göbel 1846: Cover page; Chaudoir 1850: 407;
LaFerté-Sénectěre 1851: 212, 233, 238, 263, 293; Lacordaire 1854: 213, 217,
219, 220, 221, 223, 224, 235; Chaudoir 1856: 192; Motschulsky 1860: 515; id.
1864b: 334, 347; Chaudoir 1876a: 10, 11, 12, 16; Bates 1892a: 309; Sloane 1910:
437; Andrewes 1919c: 91; id. 1923a: 58; id. 1924b: 24; id. 1930: 82; Lorenz
2005: 328.
35. Chlaenius
hamifer Chaudoir 1856
Chlaenius hamifer Chaudoir 1856: 209,
210; id. 1876: 62; Bates 1889b: 265; id. 1892b: 311; id. 1892c: 230; Bouchard
1903: 171; Lesne 1904: 69; Sloane 1910: 439; id. 1920a: 322; Andrewes 1919a:
140; id. 1924b: 24; id. 1930: 94; Lorenz 2005: 330; Löbl & Löbl 2017: 494.
= Chlaenius
bihamatus Chaudoir 1856
= Chlaenius
colombensis Jedlička 1964
= Chlaenius
queenslandicus Sloane 1910
= Dinodes bihamatus (Chaudoir
1856)
= Dinodes hamifer (Chaudoir
1856)
= Pachydinodes hamifer (Chaudoir
1856)
Specimens examined (n =
2): SJC-ZOO-CWSSMC140–141, Chinnar, 2 exs, hand picking, 26.x.2019.
Distribution: ORR - India (Kerala: Tholpetty
(Akhil 2019: 115)); SRL (Andrewes 1930: 94), MM (Andrewes 1930: 94); TAI
(Andrewes 1930: 94); PAR - BT; IN; JA; NC; HKG; NP; PA; SC; SCH (Löbl
& Löbl 2017: 494); TWN (Andrewes 1930: 94); IAR - IDS (Andrewes 1930:
94).
36. Chlaenius
nilgiricus Andrewes 1919
Chlaenius nilgiricus Andrewes 1919c: 9; id.
1930: 99; Lorenz 2005: 335.
Specimens examined (n =
3): SJC-ZOO-CWSSMC142–144, Alampetty, 2 exs, hand picking, 26.x.2019; Chinnar,
1 ex, hand picking, 26.x.2020.
Distribution: ORR - India (Tamil Nadu:
Coimbatore, Nilgiri Hills (Andrewes 1930: 99)).
Subfamily Orthogoniinae
Schaum 1857
Tribe Orthogoniini
Schaum 1857
xxi. Genus Orthogonius
Macleay 1825
Orthogonius Macleay 1825: 26; Dejean
1825: 169, 269; Schmidt-Göbel 1846: 55, 61; Lacordire 1854: 269; Walker 1858:
203; Chaudoir 1850: 434; id. 1871b: 98; Andrewes 1924b: 58; id. 1930: 245;
Csiki 1932: 1586; Jedlička 1963: 269;
Tian & Deuve 2000: 2; Lorenz 2005: 391.
= Aspectra
Schmidt-Göbel 1846
= Haplopisthius
Chaudoir 1850
= Maraga Walker
1858
37. Orthogonius
baconi Chaudoir 1871
Orthogonius baconi Chaudoir 1871d: 109;
Bates 1892a: 401; Andrewes 1930: 246; Csiki 1932: 1587; Lorenz 2005: 391; Akhil
2019: 121.
Specimens examined (n =
4): SJC-ZOO-CWSSMC145–148, Alampetty, 2 exs, hand picking, 26.x.2019; Chinnar,
2 exs, light trap, 26.x.2020.
Distribution: ORR - India (Tamil Nadu: Nilgiri
Hill; Kerala: Muthanga (Akhil 2019: 121)) MM (Andrewes 1930: 246); PAR -
India (Uttarakhand: Almora, Bengal (Andrewes 1930: 246)).
38. Orthogonius
lucidus Bates 1891
Orthogonius lucidus Bates 1891: 324–340;
Andrewes 1924b: 59; id. 1930: 248; Lorenz 2005: 392; Abhitha et al. 2009: 372.
Specimens examined (n =
8): SJC-ZOO-CWSSMC149–156, Kootar, 1ex, light trap, 26.x.2020; Alampetty, 4
exs, hand picking, 26.x.2019; Chinnar, 2exs, light trap, 26.x.2020; 1 ex, hand
picking, 26.x.2020.
Distribution: ORR - India
(Jharkhand: Chota Nagpur: Konbir, Tetara, Ranchi; Odisha: Surada; Maharashtra:
Mumbai, Igatpuri (Andrewes 1930: 248); Karnataka: Belgaum, northern Karnataka
(Andrewes 1930: 248), Bengal: Raniganj (Andrewes 1930: 248); Kerala: Kannur,
Kozhikode, Thamarassery, Wayanad: Muthanga, Idukki, Thodupuzha (Abhitha et al.
2009: 372)).
Subfamily Panagaeinae
Bonelli 1810
Tribe Panagaeini Bonelli
1810
xxii. Genus
Craspedophorus Hope 1838
Craspedophorus Hope 1838: 165;
Lacordaire 1854: 210; Chaudoir 1878: 90; Andrewes 1919a: 126; id. 1924b: 22;
id. 1930: 133; Kirschenhofer 2000: 328; Lorenz 2005: 320; Hackel &
Kirschenhofer 2014: 276; Fedorenko 2016: 2; Löbl & Löbl 2017: 638.
= Camptoderus Hope
1838
= Eudema Laporte
De Castelnau 1840
= Isotarsus LaFerté-Sénectère 1851
= Epicosmus
Chaudoir 1846
= Brachyonychus Chaudoir 1879
= Brachycosmus
Jeannel 1949
= Acanthocosmus
Jeannel 1949
39. Craspedophorus
angulatus (Fabricius 1781)
Craspedophorus angulatus (Fabricius) Andrewes
1919a: 125; id. 1921a: 154; id. 1924b: 115; id. 1924d: 462; id. 1930: 133; Jedlička 1965: 3; Kirschenhofer 2000: 323;
Baehr 2003: 446; Lorenz 2005: 320; Pang & Tian 2012: 265; Hackel & Farkac
2012: 78; Hackel & Kirschenhofer 2014: 276 & 357; Fedorenko 2016: 4;
Manthen & Hegde 2018: 206; Jithmon & Sabu 2021: 18566.
Carabus angulatus Fabricius
1781: 302; id. 1787: 197; id. 1792: 148
= Carabus angulatus
Fabricius 1781
= Pimelia fasciatus
Fabricius 1781
= Cychrus reflexus Fabricius
1801
= Panagaeus
tomentosus Vigors 1825
= Eudema bifasciatum Chaudoir
1879
= Panagaeus michardi Fairmaire
1880
= Craspedophorus
bifasciatus (Chaudoir 1879)
= Craspedophorus
fasciatus (Fabricius 1781)
= Craspedophorus
michardi (Fairmaire 1880)
= Craspedophorus
reflexus (Fabricius 1801)
= Craspedophorus
tomentosus (Vigors 1825)
= Epicosmus
bifasciatus (Chaudoir 1879)
= Eudema michardi (Fairmaire
1880)
Specimens examined (n =
2): SJC-ZOO-CWSSMC157–158, Chinnar, 2 exs, hand picking, 25.ii.2020.
Distribution: ORR - India (Andhra Pradesh;
Karnataka: Shivamoga, Mysore (Hackel & Kirschenhofer 2014: 357); Tamil
Nadu: Coimbatore (Hackel & Kirschenhofer 2014: 276 & 357); Pondicherry (Hackel & Farkac 2012: 78);
Kerala: Bonacaud (Jithmon & Sabu 2021: 18566)); SRL (Andrewes 1930: 133);
BGD (Hackel & Farkac 2012: 78); MM (Hackel & Farkac 2012: 78).
40. Craspedophorus
bifasciatus (Laporte De Castelnau 1835)
Craspedophorus
bifasciatus (Laporte De Castelnau) Andrewes 1919a: 126;
id. 1921c: 341; Andrewes 1930: 134; Kirschenhofer 2000: 323; Lorenz 2005: 320; Hackel & Farkac
2012: 78; Hackel & Kirschenhofer 2014: 276 & 346; Fedorenko 2016: 4;
Jithmon & Sabu 2021: 18567.
= Panagaeus bifasciatus Laporte De Castelnau
1835
= Epicosmus castelnaui
Chaudoir 1879
= Craspedophorus
castelnaui (Chaudoir 1879)
= Isotarsus
bifasciatus (Laporte 1835)
Distribution: ORR - India (Madhya Pradesh;
Odisha: Barkuda Island-Lake Chilka (Andrewes 1930: 134); Andhra Pradesh:
Udayagiri, Horsely Konda (Andrewes 1930: 134); Tamil Nadu: Kadayam, Coimbatore,
Bharathiyar (Jithmon & Sabu 2021: 18567), Chennai, Mahabalipuram (Hackel
& Kirschenhofer 2014: 346), Nilgiri Hills, Thiruchirapally (Andrewes 1930:
134); Pondicherry (Andrewes 1930: 134); Kerala: Chinnar (Jithmon & Sabu
2021: 18567)); SRL (Andrewes 1930: 134); BGD (Hackel & Farkac 2012: 78); MM
(Hackel & Farkac 2012: 78).
Subfamily Pterostichinae Bonelli 1810
Tribe Abacetini Chaudoir
1872
xxiii. Genus Abacetus
Dejean 1828
Abacetus Dejean 1828: 195;
Lacordaire 1854: 315; Chaudoir 1859: 126; id. 1869: 355; Tschitschérine 1898:
519, 531 & 538; id. 1902: 506; Andrewes 1924b: 44; id. 1930: 1; id. 1939:
129; Jeannel 1948: 420; Löbl & Smetana 2003: 346; Lorenz 2005: 255; Löbl
& Löbl 2017: 480.
41. Abacetus
haplosternus Chaudoir 1878
Abacetus haplosternus Chaudoir 1878: 25;
Andrewes 1930: 4; id. 1942b: 25; Lorenz 2005: 258; Divya & Sabu 2020; 9.
Specimens examined (n =
4): SJC-ZOO-CWSSMC159–162, Kootar, 3 exs, light trap, 25.x.2019; 1 ex, light
trap, 25.ii.2020.
Distribution: ORR - India (Madhya Pradesh:
Hoshangabad; Maharashtra: Nagpur (Andrewes 1930: 4)); TAI (Andrewes 1930: 4); PAR
- India (Himachal Pradesh: Katrain; Uttarakhand: Almora, Ranikhet, Haldwani
(Andrewes 1930: 4)); IAR - IDS (Andrewes 1930: 4).
xxiv. Genus
Cosmodiscus Sloane 1907
Cosmodiscus Sloane 1907: 371;
Andrewes 1920b: 445; id. 1930: 131; Löbl & Smetana 2003: 443; Lorenz 2005:
260; Kushwaha & Hegde 2015: 396, 401; Löbl & Löbl 2017: 481.
42. Cosmodiscus
picturatus Andrewes 1920
Cosmodiscus picturatus Andrewes 1920b: 447;
id. 1921c: 345; id. 1930: 131; Lorenz 2005: 260; Kushwaha & Hegde 2015:
396, 401; Divya & Sabu 2020: 11.
Specimens examined (n =
2): SJC-ZOO-CWSSMC163–164, Alampetty, 2 exs, light trap, 26.x.2019.
Distribution: ORR - India (Uttar Pradesh:
Fyzabad, Odisha: Rambha: Ganjam, Barkuda and Gopkuda Island, lake Chilka;
Maharashtra: Nagpur; Andhra Pradesh: Jammelamadugu (Andrewes 1930: 131);
Kerala: Kozhikode (Divya & Sabu 2020: 11)).
Tribe Cratocerini
Lacordaire 1854
xxv. Genus Caelostomus MacLeay 1825
Caelostomus MacLeay 1825: 23; Andrewes 1924b:
44; id. 1930: 55; Jeannel 1948: 383; Löbl I & Smetana 2003: 471; Lorenz
2005: 249; Faisal & Singh 2014: 342; Löbl
& Löbl 2017: 678.
* 43. Caelostomus
sculptipennis (Motschulsky 1859)
Caelostomus
sculptipennis (Motschulsky) Chaudoir 1872c: 13; Tschitschérine 1900b:
263 (note); Andrewes 1928: 22; id. 1930: 57; Straneo 1938: 56; Lorenz 2005:
250; Divya & Sabu 2020: 12.
= Stomonaxus
sculptipennis Motschulsky 1859
= Stomonaxus
sculpticollis Motschulsky 1859
= Caelostomus
sculpticollis (Motschulsky 1859)
Specimens examined (n =
1): SJC-ZOO-CWSSMC165, Chinnar, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India
(Tamil Nadu: Nilgiri Hills (Straneo 1938: 56)); SRL (Andrewes 1930: 57).
Tribe Pterostichini
Bonelli 1810
xxvi. Genus
Trigonotoma Dejean 1828
Trigonotoma Dejean 1828: 182; Brulle
1834: 333; Chaudoir 1852: 71; Lacordaire 1854: 311; Chaudoir 1868: 158;
Tschitschérine 1900b: 180; Kuntzen 1911: 182; id. 1914: 60; Andrewes 1930: 352;
id. 1939: 138; Saha & Halder 2000: 20; Löbl & Smetana 2003: 520; Lorenz
2005: 300; Dubault et al. 2008: 240; Kushwaha & Hegde 2015: 396, 401; Löbl
& Löbl 2017: 755.
#44. Trigonotoma
oberthueri Tschitschérine 1894
Trigonotoma oberthueri Tschitschérine 1894b:
444; Kuntzen 1914: 63; Andrewes 1930: 355; Löbl & Smetana 2003: 520; Lorenz
2005: 300; Löbl & Löbl 2017: 755; Divya & Sabu 2020: 22.
Specimens examined (n =
1): SJC-ZOO-CWSSMC166, Chinnar, 1 ex, hand picking, 26.x.2019.
Distribution: PAR - India (West Bengal:
Pedong, Gopaldhara, Mungphu, Kurseong, Lebong (Andrewes 1930: 355)).
Subfamily Scaritinae
Bonelli 1810
Tribe Clivinini
Rafinasque 1815
xxvii. Genus Clivina
Latreille 1802
Clivina Latreille 1802: 96;
Bonelli 1813: 480; Dejean 1825: 411; Schmidt-Göbel 1846 (cover); Motschulsky
1861: 101; Putzeys 1863: 29 & 68; id. 1867: 94; id. 1868: 10; id. 1873: 15;
Fleisch 1899: 33; Tschitschérine 1904: 258; Andrewes 1919b: 470; id. 1924b: 11;
id. 1926c: 372; id. 1929: 344, 351; id. 1930: 110; Balkenohl 2001: 13; Lorenz
2005: 141.
45. Clivina brevior Putzeys
1866
Clivina brevior Putzeys 1866: 126; Bates
1892a: 277; Andrewes 1926c: 375; id. 1929: 355, 378; id. 1930: 112;
Balkenohl 2001: 14; Lorenz 2005: 142; Abhitha 2010: 105.
Specimens examined (n =
1): SJC-ZOO-CWSSMC167, Chinnar, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (New Delhi: Pusa
(Andrewes 1930: 112); Kerala: Kozhikode: Kuttikattoor, Medical College,
Thamarassery (Abhitha 2010: 105)); MM (Andrewes 1930: 112); IAR - MLS
(Andrewes 1930: 112).
46. Clivina lobata
Bonelli 1813
Clivina lobata Bonelli 1813: 481;
Dejean 1825: 414; Putzeys 1861: 50; id. 1867: 121, 122, 125; id. 1868: 1, 8;
Bates 1892a: 276; Andrewes 1919a: 209; id. 1921c: 340; id. 1922: 392; id.
1924b: 11, 462; id. 1926c: 875; id. 1929: 355, 375; id. 1930: 114; Lorenz 2005:
143; Abhitha 2010: 107; Löbl & Löbl 2017: 255.
Specimens examined (n =
1): SJC-ZOO-CWSSMC168, Kootar, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Kerala:
Kozhikode: Thamarassery, Wayanad: Thirunelli (Abhitha 2010: 107)); MM (Andrewes
1930: 114); TAI (Andrewes 1930: 114); PAR - JA (Löbl & Löbl 2017:
255).
xxviii. Genus Pseudoclivina
Kult 1947
Pseudoclivina Kult 1947: 30; id. 1951:
18; Balkenohl 2001: 18; Lorenz 2005: 145; Löbl & Löbl 2017: 258.
*47. Pseudoclivina
costata (Andrewes 1929)
Pseudoclivina costata (Andrewes) 1929: 354,
364; id. 1930: 113; Kult 1951: 18; Bakenohl 2001: 18; Lorenz 2005: 145.
= Clivina costata Andrewes 1929: 354
Specimens examined (n =
1): SJC-ZOO-CWSSMC169, Alampetty, 1 ex, light trap, 25.ii.2020.
Distribution: ORR - India (Tamil Nadu: Nilgiri
Hills (Andrewes 1930: 113)).
48. Pseudoclivina
memnonia (Dejean 1831)
Pseudoclivina memnonia (Dejean) Kult 1947: 30;
id. 1951: 18; Balkenohl 2001: 19; Lorenz 2005: 145; Abhitha 2010: 108; Löbl
& Löbl 2017: 259.
Clivina memnonia Dejean 1831: 503; Putzeys
1846: 588; Bouchard 1903: 169; Andrewes 1919a: 187, 206; id. 1924b: 115; id.
1926c: 373; id. 1927: 105; id. 1929: 354, 362; id. 1930: 115; Saha & Biswas
1985: 120.
= Clivina memnonia
Dejean 1831
= Clivina indica
Putzeys 1846
= Clivina rugosifrons
Nietner 1856
= Clivina recta
Walker 1858
= Pseudoclivina
indica (Putzeys 1846)
= Pseudoclivina recta (Walker
1858)
= Pseudoclivina
rugosifrons (Nietner 1856)
Specimens examined (n =
2): SJC-ZOO-CWSSMC170–171, Alampetty, 1 ex, light trap, 26.x.2019; Chinnar, 1
ex, light trap, 26.x.2019.
Distribution: ORR - India (Kerala: Idukki:
Chinnar; Kozhikode: Thamarassery, Engapuzha; Kasargod: Periya; Wayanad: Sulthan
Bathery, Ambalavayal, Panamaram, Thirunelli, Muthanga, Tholpetty (Abhitha 2010:
108)); SRL (Andrewes 1930:115); MM (Andrewes 1930:115); PAR - GUA, HAI,
YUN (Löbl & Löbl 2017: 259); IAR - IDS (Andrewes 1930:115).
Tribe Dyschiriini W.
Kolbe 1880
xxix. Genus Dyschirius
Bonelli 1810
Dyschirius Bonelli 1810: Panzer
1813: 67; Stephens 1827: 37, 40; Putzeys 1846: 524; Lacordaire 1854: 202;
Putzeys 1867: 32; Fleischer 1899: 8; Andrewes 1919: 99; Müller 1922: 33;
Andrewes 1926c: 377; id. 1929: 390; id. 1930: 159; Jeannel 1941: 250, 260, 275;
id. 1946: 213, 215, 218; Moore &
Brown 1979: 123; Clopton 1991: 53, 59; Saha et al. 1992: 9; Balkenohl
1994: 27; Fedorenko 1996: 5, 9, 11; Lorenz 2005: 151; Bulirsch 2009: 559; id.
2011: 1; Bousquet 2012: 431; Allegro & Bulirsch 2012: 235; Hogan 2012: 106,
111, 116, 231; Kushwaha & Hegde 2015: 399, 419; Fedorenko 2016: 439; Ghannem
et al. 2016: 69; Bulirsch & Stachowiak 2017: 137; Löbl & Löbl 2017:
263; Bulirsch 2018: 229.
49. Dyschirius paucipunctus Andrewes 1929
Dyschiriodes
paucipunctus (Andrewes) Lorenz 2005: 154.
Dyschirius mahratta Var. paucipunctus
Andrewes 1929: 392, 397; id. 1930:160.
= Dyschiriodes
paucipunctus (Andrewes 1929)
Specimens examined (n =
3): SJC-ZOO-CWSSMC172–174, Kootar, 3 exs, light trap, 26.x.2019.
Distribution: ORR - India (Maharashtra:
Pune; Karnataka: Belgaum (Andrewes 1930: 160)); SRL (Andrewes 1930: 160).
Tribe Scaritini Bonelli
1810
xxx. Genus Oxylobus
Chaudoir 1855
Oxylobus Chaudoir 1855: 5; id.
1879: 129; Andrewes 1924b: 8; id. 1929: 292; id. 1930: 252; Lorenz 2005: 141.
50. Oxylobus
asperulus Chaudoir 1857
Oxylobus asperulus Chaudoir 1857: 58; id.
1879: 133; Andrewes 1922: 215; id. 1924b: 129; id. 1929: 296, 311. id. 1930:
252; Lorenz 2005: 141.
Specimens examined (n =
1): SJC-ZOO-CWSSMC175, Alampetty, 1 ex, hand picking, 26.x.2019.
Distribution: ORR - India (Andhra
Pradesh: Chittur district, Horseley Konda; Karnataka: Mysore; Tamil Nadu: Pillur,
Kodaikanal, Yercaud, Madura, Nilgiri Hills, Shembaganur; Kerala: Dhoni forest,
southern Malabar (Andrewes 1930: 252)); SRL (Andrewes 1930: 252).
# ssp. Oxylobus
asperulus amyntas Andrewes 1924
Oxylobus amyntas Andrewes
1924b: 70; id. 1929: 296, 313. id. 1930: 252; Lorenz 2005: 141.
Specimens examined (n =
2): SJC-ZOO-CWSSMC176–177, Alampetty, 2 exs, hand picking, 26.x.2019.
Distribution: ORR - India (Madhya Pradesh:
Majgaon, Motinala, Mukhi (Andrewes 1930: 252)).
51. Oxylobus porcatus
(Fabricius 1798)
Oxylobus porcatus (Fabricius)
Heyne-Taschenberg 1894: 3: 32; id. 1895: 20; Andrewes 1921a: 157; id. 1924b: 8;
id. 1929: 295, 305; Andrewes 1930: 254; Lorenz 2005: 141.
Scarites porcatus Fabricius 1798: 43;
Hope 1838: 95; Motschulsky 1855: 40.
= Scarites porcatus
Fabricius 1798
= Oxylobus costatus
Chaudoir 1879
= Oxylobus minor Tschitschérine
1894a
= Oxylobus
obliterates Andrewes 1929
Specimens examined (n =
3): SJC-ZOO-CWSSMC178–180, Alampetty, 3 exs, hand picking, 26.x.2019.
Distribution: ORR - India (Punjab:
Baddia; West Bengal: Sahibganj, Rajmahal, Giridih; Jharkhand: Chakardharapore, Konbir, Chota Nagpur-
Tetara, Tinpahar; Madhya Pradesh: Jubbulpore, Majgaon, Motinala; Chhattisgarh:
Chitrakot; Odisha: Barkuda Island, Barkul, Chilka lake; Andra Pradesh:
Visakhapatnam, Chittoor, Horseley Konda; Karnataka: Belgaum; Tamil Nadu: Coimbatore, Nilgiri
Hills, Shevaroy Hills, Madura, Palni Hills, Kallar, Pillur, Ootacamund,
Shembagannur; Kerala: Malabar Coast (Andrewes 1930: 254)); SRL (Andrewes 1930:
254).
Subfamily Trechinae
Bonelli 1810
Tribe Bembidiini
Stephens 1827
xxxi. Genus Elaphropus
Motschulsky 1839
Elaphropus Motschulsky 1839: 73;
Erwin 1975: 1; Kopecky 2002: 63; Lorenz 2005: 207; Löbl & Löbl 2017: 342.
* 52. Elaphropus nigellus
(Andrewes 1935)
Elaphropus nigellus (Andrewes)
Lorenz 2005: 210.
= Tachys nigellus Andrewes 1935
= Tachyura nigella (Andrewes
1935)
Specimens examined (n =
21): SJC-ZOO-CWSSMC181–201, Chinnar, 2 exs, light trap, 26.x.2019; Alampetty, 7
exs, light trap, 26.x.2019; 2 exs, pitfall trap, 26.x.2019; 2 exs, hand
picking, 26.x.2019; 1 ex, light trap, 25.ii.2020; 2 exs, pitfall trap,
25.ii.2020; 2 exs, hand picking, 25.ii.2020; Kootar, 2 exs, light trap,
26.x.2019; 1 ex, hand picking, 26.x.2019.
Distribution: ORR - India (Tamil Nadu:
Chennai, Nilgiri Hills; Kerala: Nilambur (Andrewes 1935: 277)).
* 53. Elaphropus nilgiricus
(Andrewes 1925)
Elaphropus nilgiricus
(Andrewes) Lorenz 2005: 210.
Tachys nilgiricus Andrewes 1925: 446; id.
1930: 334; id. 1935: 265.
= Tachys nilgiricus Andrewes 1925
= Tachys unisculptus
Andrewes 1925
= Elaphropus
unisculptus (Andrewes 1925)
= Tachyura nilgirica (Andrewes
1925)
Specimens examined (n =
2): SJC-ZOO-CWSSMC202–203, Alampetty, 1 ex, light trap, 26.x.2019; 1 ex, light
trap, 25.ii.2020.
Distribution: ORR - India (Karnataka:
Mysore (Andrewes 1930: 334); Tamil Nadu: Nilgiri Hills (Andrewes 1935:
446 )); SRL (Andrewes 1930: 334).
# 54. Elaphropus
politus (Motschulsky 1851)
Elaphropus politus (Motschulsky) Lorenz
2005: 210; Kushwaha & Hegde 2015: 395.
Tachys politus Motschulsky 1851: 509;
Putzeys 1875b: 743; Bouchard 1903: 170; Andrewes 1919a: 199; id. 1921a: 146;
id. 1925: 448; id. 1930: 338; id. 1935: 269.
= Tachys politus Motschulsky 1851
= Tachyura polita (Motschulsky
1851)
Specimens examined (n =
20): SJC-ZOO-CWSSMC204–223, Chinnar, 2 exs, light trap, 26.x.2019; Alampetty, 5
exs, light trap, 26.x.2019; 2 exs, pitfall trap, 26.x.2019; 3 exs, hand
picking, 26.x.2019; 2 exs, light trap, 25.ii.2020; 1 ex, pitfall trap,
25.ii.2020; Kootar, 4 exs, light trap, 25.x.2019; 1 ex, pitfall trap,
25.ii.2020.
Distribution: ORR - India (Uttar Pradesh:
Auraiya, Fatehpur, Muradganj, Mathura, Kishori Kunj, Jhansi, Shahjahanpur (Kushwaha & Hegde
2015: 395)); SEA (Andrewes 1935: 448).
DISCUSSION
This is the first report
about ground beetles from a natural habitat in the eastern slopes of Western
Ghats and it represents the carabid composition in a dry deciduous forest in
the southern WGs. Fifty-four species belonging to 11 subfamilies (Harpalinae:
15 species, Lebiinae: 14, Scaritinae: 7, Pterostichinae: 4, Anthiinae: 3,
Trechinae: 3, Licininae: 2, Orthogoniinae: 2, Panagaeinae: 2, Brachininae: 1,
Dryptinae: 1), and 31 genera were recorded. Harpalinae, Lebiinae, and Scaritinae
are the species-rich subfamilies with 15, 14, and seven species respectively,
in the study region which is a representative of the dry forest habitat in the
rain shadow slopes of the southern WG. Two species—Stenolophus lucidus (Harpalinae) and Amblystomus aenescens (Harpalinae)—are
first records from India (Image 1A,B). Four species, Stenolophus bajaurae (Harpalinae), Amblystomus indicus (Harpalinae), Trigonotoma
oberthueri (Pterostichinae), and Elaphropus
politus (Trechinae) (Image 2I,A,J,E) are first reports from southern India
and Oxylobus asperulus amyntas (Scaritinae) is the first record of the
subspecies from southern India (Image 2G). Amblystomus indicus was reported earlier from
Sri Lanka and eastern & western India (Bates 1886, 1892; Andrewes 1930) and
the record in southern India is significant indicating its continuous
distribution in Sri Lanka and southern India. Trigonotoma oberthueri, a
species with earlier reports only from the PAR in the central and eastern
Himalayan region (Andrewes 1930; Löbl & Löbl 2017) is recorded from the
Oriental region. Six species (Macrocheilus chinnarensis (Anthiinae), Ophoniscus puneensis (Harpalinae), Caelostomus
sculptipennis (Pterostichinae), Pseudoclivina costata (Scaritinae),
Elaphropus nigellus (Trechinae), E. nilgiricus (Trechinae)
(Image 2F,B,H,C,D) are endemic to the WG and Sri Lanka biodiversity hot spot. Macrocheilus
chinnarensis is a recently discovered new local endemic
species (Akhil et al. 2019). Ophoniscus puneensis
is recorded for the first time from south WG after its
discovery in the northern WG (Kataev 2018). Pseudoclivina costata and Elaphropus
nigellus are endemic to the southern WG (Andrewes 1925, 1929,
1930, 1935) and it is the first record of the species from the eastern slopes
of the WG. Caelostomus sculptipennis and Elaphropus nilgiricus are
known only from southern WG and Sri Lanka (Andrewes
1925, 1928, 1930, 1935; Straneo 1938; Divya & Sabu 2020).
For
figure & images - - click here
REFERENCES
Abhitha, P. (2010). Forest litter faunal
diversity and abundance in relation to litter chemical quality and systematics
of Carabid beetles. PhD Thesis. Forest Research Institute University Dehra Dun,
Uttarakhand, 176 pp (Unpublished).
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