Journal of Threatened Taxa |
www.threatenedtaxa.org | 26 June 2021 | 13(7): 18687–18694
ISSN 0974-7907 (Online) | ISSN 0974-7893
(Print)
https://doi.org/10.11609/jott.7207.13.7.18687-18694
#7207 | Received 22 February 2021 | Final
received 27 May 2021 | Finally accepted 30 May 2021
Population assessment of the
endangered Western Hoolock Gibbon Hoolock hoolock
Harlan, 1834 at Sheikh Jamal Inani National Park,
Bangladesh, and conservation significance of this site for threatened wildlife
species
M. Tarik Kabir 1, M.
Farid Ahsan 2, Susan M. Cheyne 3, Shahrul
Anuar Mohd Sah 4, Susan Lappan
5, Thad Q. Bartlett 6 &
Nadine Ruppert 7
1,4,5,7 School of Biological Sciences, Universiti Sains Malaysia, 11800 Pulau Pinang, Malaysia.
2 Department of Zoology, University
of Chittagong, Chattogram, Bangladesh.
3 Department of Social Science,
Oxford Brookes University, Oxford, Oxfordshire, GB, United Kingdom.
5 Department of Anthropology,
Appalachian State University, Boone, NC, United States of America.
6 Department of Anthropology, The
University of Texas at San Antonio, 6900 North Loop 1604 West, San Antonio, TX
78249, United States of America.
1,2,3,5,6,7 IUCN SSC PSG Section on Small
Apes.
1 tarikkabir84@gmail.com, 2 faridahsan55@yahoo.com,
3 section.small.apes@gmail.com, 4 sanuar@usm.my, 5 lappansm@appstate.edu,
6 thad.bartlett@utsa.edu, 7 n.ruppert@usm.my
(corresponding author)
Editor: Honnavalli N. Kumara, Salim Ali Centre for Ornithology and Natural
History, Coimbatore, India.
Date of
publication: 26
June 2021 (online & print)
Citation: Kabir, M.T., M.F. Ahsan, S.M. Cheyne, S.A.M. Sah, S. Lappan, T.Q. Bartlett
& N. Ruppert (2021). Population assessment
of the endangered Western Hoolock Gibbon Hoolock hoolock
Harlan, 1834 at Sheikh Jamal Inani National Park,
Bangladesh, and conservation significance of this site for threatened wildlife
species. Journal of Threatened Taxa 13(7): 18687–18694. https://doi.org/10.11609/jott.7207.13.7.18687-18694
Copyright: © Kabir et al. 2021. Creative Commons Attribution 4.0 International
License. JoTT
allows unrestricted use, reproduction, and distribution of this article in any
medium by providing adequate credit to the author(s) and the source of
publication.
Funding: This study was funded by the IUCN Section on Small Apes, Gibbon Conservation Alliance, Wildlife
Conservation Network, International Primatological Society as the Galante Family Winery Conservation Scholarship, Global Wildlife Conservation: Primate Action Fund (Grant Number: 5385.006-0318) and Rufford
Foundation Small Grant (Grant Number: 33003-01) through the Malaysian Primatological Society.
Competing interests: The authors declare no competing interests.
Author
details: M. Tarik Kabir is PhD
candidate of Universiti Sains
Malaysia and a wildlife biologist in Bangladesh. His major interests are
primate conservation, community-based wildlife conservation, and social
behavior and ecology of gibbons. Dr. M. Farid Ahsan, Professor of Zoology
at University of Chittagong, Bangladesh, member of the IUCN Section on Small
Apes, and a Fellow of the Zoological Society of Bangladesh. He teaches
undergraduates and graduates courses in wildlife biology and conservation. His
interest is in primate behaviour, ecology, and
biodiversity conservation, and creating awareness about snakebites. Dr.
Susan M. Cheyne is the Vice-Chair of the IUCN SSC PSG Section on Small
Apes and a Teaching Fellow at Oxford Brookes University. Her interests are in
small ape conservation in the wild, in captivity and in creating opportunities
for these apes to return to the wild following capture to the illegal pet
trade. Dr.
Shahrul Anuar Mohd Sah is a Professor of
Zoology at Universiti Sains
Malaysia. He has a long experience in teaching wildlife conservation and
management, wildlife ecology, and animal behaviour.
His research interests are particularly in the field of vertebrate biology,
ecology and behaviour, and mainly focused on the
population studies of small mammals, birds and herpetofauna. Dr.
Susan Lappan is a behavioural
ecologist with a primary interest in primate conservation. She is an Associate
Professor in the College of Arts and Sciences at Appalachian State University,
USA, and a scientific advisor for Malaysian Primatological
Society. Dr.
Thad Q. Bartlett is a member of the IUCN Section on Small Apes and
Professor of Anthropology at The University of Texas at San Antonio where he
teaches undergraduate and graduate courses in human nature, human origins, and
primate behavior and ecology. His primary research focus is on gibbon socioecology. Dr. Nadine Ruppert
is a zoologist focusing on primates, human-wildlife interactions, plant-animal
interactions, and conservation issues related to Malaysia. She is a senior lecturer
at Universiti Sains
Malaysia, vice president of the Malaysian Primatological
Society and member of the IUCN Section on Small Apes.
Author
contributions: Research methodology and first manuscript draft: Kabir, M.T. and Ruppert, N.; field work: Kabir, M.T.; advisory role during
field work and manuscript corrections: Ahsan, M.F., Cheyne, S.M., Sah, S.A.M., Lappan, S. and
Bartlett, T.Q.
Acknowledgements:
We gratefully acknowledge the financial support from IUCN Section on
Small Apes, Gibbon Conservation Alliance, Wildlife Conservation Network,
International Primatological Society, Rufford Foundation Small Grant (Grant Number: 33003-01)
through the Malaysian Primatological Society, and
Global Wildlife Conservation: Primate Action Fund (Grant Number: 5585.006-0318)
for the field work. We also acknowledge the Bangladesh Forest Department; Ishtiaq Uddin Ahmad, former Chief Conservator of Forests,
Bangladesh Forest Department; Divisional Forest Officer, Cox’s Bazar South
Forest Division; local forest officials, and our local research assistants for
their generous support and suggestions in support of this effort. We also
acknowledge to School of Biological Sciences, Universiti
Sains Malaysia, Malaysia for providing us necessary
supports. We are also grateful to Sayam U. Chowdhury, an ornithologist, for identification of
Slaty-backed Flycatcher. We gratefully acknowledge
the participants of our focus group discussions.
Abstract: Sheikh Jamal Inani
National Park (Inani) is a wildlife habitat in
Bangladesh located under the Cox’s Bazar South Forest Division, Cox’s Bazar. It
constitutes significant habitat for the charismatic and globally ‘Endangered’
Western Hoolock Gibbon Hoolock hoolock in
Bangladesh. Here, we show that Inani is a
poorly-known gibbon habitat with a population of seven groups, comprising a
total of 18 individuals. Among them, 77.8% were adults (males and females), and
11.1%, 5.6%, and 5.6% were sub-adults, juveniles, and infants, respectively,
indicating low reproductive output. Five of seven groups had no offspring
present in the group, and the mean group size of 2.57 individuals/group is low
compared to other habitats in Bangladesh. Beside Western Hoolock Gibbon, Inani is home to many threatened wildlife species. The
first record of the Slaty-backed Flycatcher Ficedula erithacus
in Bangladesh occurred in Inani, adding this new
species to the national bird checklist of Bangladesh. The presence of the
globally ‘Endangered’ Asian Elephant Elephas maximus, Phayre’s Langur Trachypithecus
phayrei, & Elongated Tortoise Indotestudo elongata
and the globally ‘Vulnerable’ Northern Pig-tailed Macaque Macaca
leonina, Capped Langur Trachypithecus
pileatus, Indian Leopard Panthera
pardus, & Asiatic Softshell Turtle Amyda cartilaginea
highlight the importance of Inani as a conservation
area in Bangladesh. The Western Hoolock Gibbon and other threatened wildlife of
Inani are now on the verge of local extinction due to
a sharp increase in forest resource extraction resulting from the recent influx
of large numbers of Rohingya refugees from Myanmar, many of whom have settled
around Inani. Through stakeholder interviews in the
area, we have identified feasible and measurable conservation actions at Inani that are urgently needed to prevent further loss of
wildlife and to protect this important gibbon habitat.
Keywords: Cox’s Bazar South Forest
Division, Rohingya, Slaty-backed Flycatcher.
INTRODUCTION
Sheikh Jamal Inani National Park (short: Inani) is the southern-most natural, although heavily
degraded, forest in Bangladesh. It was previously known as Inani
Reserved Forest, and was declared a National Park in 2019 by the Bangladesh
Government. Inani includes the last remnants of
degraded natural forest in Cox’s Bazar South Forest Division and supports many
globally threatened wildlife species (Kabir et al. 2014, 2015, 2017). These
forests also form a wildlife corridor between Myanmar and Bangladesh that is
recognized in Bangladesh as a prominent Asian Elephant corridor (IUCN
Bangladesh 2018).
Rohingya
refugees are defined by the People’s Republic of Bangladesh as
‘forcibly-displaced Myanmar nationals’ (UNDP Bangladesh and UN WOMEN Bangladesh
2018). About a million Rohingya refugees have settled in Bangladesh in
successive waves of displacement since the early 1990s (https://www.unhcr.org/rohingya-emergency.html),
of which 716,915 are new arrivals since 25 August 2017 (https://data2.unhcr.org/en/situations/myanmar_refugees). They have settled at Ukhia
and Teknaf upazila (sub
district) under the Cox’s Bazar District of Bangladesh. The majority of them
have settled around or inside the Ukhia Reserved
Forest, Sheikh Jamal Inani National Park, and Teknaf Wildlife Sanctuary, administered by the Bangladesh
Government and UNHCR. Makeshift camps and fuel-wood collection have had
significant impacts on forested areas, resulting in forest degradation and
habitat loss, wildlife habitat fragmentation, loss of wildlife corridors, and
an increase in elephant-human conflict (UNDP Bangladesh and UN WOMEN Bangladesh
2018).
Several recent publications over the last 12 years describe wildlife
diversity and conservation in Inani (e.g., Akhter et
al. 2009; Ahmed et al. 2011; Rahman & Mannan 2011; Kabir et al. 2014, 2015,
2017; Haidar et al. 2017). Drastically decreasing habitat quality at Inani due to forest loss and other threats, such as
encroachment and extraction of forest products by nearby local and Rohingya
communities, however, are driving the Western Hoolock Gibbon, as well as other
wildlife species, to the brink of local extinction.
No recent information has been published on the population status of
Western Hoolock Gibbon Hoolock hoolock Harlan,
1834 at this site (Image 1). New information is provided in this paper on the
population status of Western Hoolock Gibbons at Inani,
and we report the occurrence of other globally threatened wildlife species,
indicating the value of the site. Through stakeholder interviews in the area,
we identified in situ conservation initiatives that should be undertaken
immediately to protect Western Hoolock Gibbon and other threatened species at Inani.
MATERIALS AND METHODS
Study
Area
The Sheikh Jamal Inani National Park (21.226642
N and 092.081416 E) covers an area of 7085.16 ha of hill forest in the Inani Forest Range under the Cox’s Bazar South Forest
Division of Cox’s Bazar District of Bangladesh. It is bordered by the Himchhari National Park in the north, Teknaf
Wildlife Sanctuary in the south, Ukhia Reserved
Forest in the east, and the Bay of Bengal in the west.
The vegetation of Inani is mixed-evergreen
forest dominated by degraded secondary forests. Major tree species are Garjan Dipterocarpus spp.,
Chapalish Artocarpus
chama, Chundul Tetrameles nudiflora,
Civit Swintonia
floribunda, Telsur Hopea
odorata, Shimul Bombax
spp., Pitraj Aphanamixis
polystachya, Koroi Albizia spp., Bandorholla
Duabhanga grandiflora, Jam Syzygium spp., Rata Amoora
wallichii, Nageshwar Mesua ferrea,
Uri-am Magnifera longipes,
Bhadi/Jiol Lannea coromandelica,
Jarul Lagerstroemia spp., Gamar Gmelia arborea,
Figs Ficus spp., and Ajuli Dillenia pentagyna (Kabir 2012).
The composition of the undergrowth, including bamboos, varies
considerably from place to place. The most common species are Mulibansh Melocanna bambusoides, Mitinga Bambusa tulda,
Ground Orchid Geodorum spp., Galla Bet Daemonorops jenkinsianus, and Bet Calamus spp.. There is an
abundance of creepers, lianas, and epiphytes, including Tinospora
cordifolia, Vitis
spp., Spatholobus roxburghii,
Entada pursaetha,
Derris spp., Ipomoea spp., Passiflora
spp., Oberonia spp., and others.
Methods
Western
Hoolock Gibbon habitats in Bangladesh consist only of small habitat fragments,
in contrast to the larger, more continuous habitats of the species in other
countries (Ahsan 1994; Geissmann et al. 2013; Ray et
al. 2015). A gibbon population census was conducted by the total-count method
and groups were detected at established listening posts (following Brockelman
& Ali 1987; Cheyne et al. 2007; Brockelman et al. 2009). One observer sat
at one listening post carefully noting the singing times and durations of
singing bouts of gibbon pairs, taking compass bearings, and estimating the
distance from the singing pair to the listening post. Upon visual encounters,
observer(s) assessed the group composition.
Adult males, adult females, subadult males, subadult females, juveniles,
and infants were estimated on the basis of the body size and coat colour (Kakati et al. 2009), and behavioral
pattern (Ahsan 1994). Groups were distinguished by location, group composition
and distance between groups, and all groups identified were given a distinct
identification number for long-term monitoring. Gibbon groups were monitored
from January 2017 to January 2021 to confirm group compositions. Gibbon
population monitoring was conducted from early morning to early afternoon (0600
to 1400 h) for a period of four consecutive days/month from October to April
during the monitoring period. The occurrence of other threatened wildlife
species was confirmed opportunistically through direct visual observations
during field trips from January 2013 to January 2021.
Threat
assessment was conducted through direct field observations and feasible
conservation measures were identified in discussions with focus groups,
including forest-dependent people, nearby communities and villagers, community
patrol groups, local community leaders and other relevant stakeholders, such as
forest department staff (BOBLME 2013; Alam et al.
2014). Three focus-group discussions (FGD) were conducted with the participants
at Boro Inani, Patuartake, and Swankhali between
March and June 2018. There were 10–12 participants in each FGD. Participants
were selected in consultation with the local forest department and village
headmen. Predefined questionnaires were completed to assess the perceived
impact of the huge Rohingya influx to Inani and to
identify possible conservation measures to save the wildlife at Inani, including its Western Hoolock Gibbons (Alam et al. 2014).
RESULTS
Seven groups
of Western Hoolock Gibbons consisting of 18 individuals were confirmed to reside
in Sheikh Jamal Inani National Park during the study
period (Table 1). Six groups were reported from Inani
Forest Beat (local administration unit of Bangladesh Forest Department) and one
from Swankhali Forest Beat (Inani
Forest Range). Only two of these groups (Groups 3 and 4) showed evidence of
reproduction during the study period, including an adult pair with a subadult
and an infant, and an adult pair with a subadult and a juvenile (Table 1). The
mean group size was 2.57 individuals (n= 7). Synchronous singing by Groups 1,
2, 3, & 4 was heard at least twice, which indicates that they were separate
groups. The area inhabited by Groups 1–4 is considered as the core area for
immediate conservation action (Figure 1). Groups 1, 2, 3, 5, 6, & 7 were
first observed in 2014 during an opportunistic wildlife survey at Inani and at that time, each group consisted of only an
adult male and female. Group 3 produced an offspring in early 2015 and again in
January 2021.
Inani is a significant habitat for many globally threatened wildlife species,
which also require immediate conservation initiatives. The presence of the
Indian Leopard Panthera pardus
fusca in Cox’s Bazar District of Bangladesh was
first confirmed in 2014 in the core gibbon habitat of Inani
(Kabir et al. 2017), with additional sightings in 2017 and 2018 (M. Tarik
Kabir, pers. obs. 2017 & 2018). During the gibbon surveys, we also made the
first observations of the Slaty-backed Flycatcher Ficedula erithacus
in Bangladesh (Image 2). It was identified by its orange underparts, deep blue
upperparts and black tail with white base in males (Image 2). This species was
previously reported as having a global distribution in Bhutan, China, India,
Laos, Myanmar, Nepal, Vietnam, and Thailand (BirdLife
International 2016) and now we confirm that its range extends into Bangladesh.
It was first sighted in January 2014 in an area dominated by shrubs and
homestead vegetation near human habitations and was sighted again at the same
place in February 2016.
The globally
‘Endangered’ Asian Elephant Elephas maximus, Phayre’s
Langur Trachypithecus phayrei,
& Elongated Tortoise Indotestudo elongata and the ‘Vulnerable’ Capped Langur Trachypithecus pileatus,
Northern Pig-tailed Macaque Macaca leonina, Indian Leopard Panthera
pardus, & Asiatic Softshell Turtle Amyda cartilaginea
were also observed in the core gibbon habitat of Inani.
IUCN
Bangladesh (2018) has estimated that the total population of elephants in the
five forest ranges of the Cox’s Bazar South Forest Division includes only 38
individuals (31–45). Elephants are now isolated in Inani,
Ukhia, Shilkhali, Whykheong, and Teknaf forest
ranges due to the blocking of the Ukhia-Ghundhum
Elephant Corridor by Rohingya settlements since 2017 (Irfanullah
2018). Focus-group discussions with the relevant stakeholders showed that
elephant-human interaction has dramatically increased at Inani,
in the area of Mohammad Shafir Beel,
after the recent influx of Rohingya refugees who live around the forest and
collect firewood and other forest resources on an unsustainable level. The
interviewees also agreed that the wildlife habitat of Inani
will vanish in a short period of time if the current situation is not
mitigated. Beside the negative impacts on wildlife and habitat quality
resulting from the influx of refugees, the gibbon habitat in Inani has also been destroyed and degraded by illegal
resource harvesting and encroachment by local communities and forest-dependent
people. Local communities collect the stems of saplings of various tree species
and use them as poles for their betel-leaf vineyards. The interviewees stated
that they think that habitat destruction and degradation may be mitigated
through regular patrolling of the forest department and with direct involvement
of the local community, more dialogue among policy makers and the forest-living
people, and an extensive habitat restoration programme.
DISCUSSION
The Western
Hoolock Gibbon is a ‘Critically Endangered’ species in Bangladesh (IUCN
Bangladesh 2015) and an ‘Endangered’ species globally (Brockelman et al. 2019).
About 282 individuals were reported in Bangladesh in surveys over a decade ago
(Islam et al. 2006). Islam et al. (2006) observed two groups of gibbons in the Inani Range and five in the Ukhia
Range during eight-day (Inani) and nine-day (Ukhia) survey periods. Based on our survey results, it
seems likely that Inani supported a larger gibbon
population during the 2003–2004 survey period, and that not all gibbon groups
were observed within the short survey period. Moreover, at that time, the
habitat quality was much better than presently, but gibbons have now become
locally extinct in Ukhia (M. Tarik Kabir, pers. obs.
2020).
It was
revealed in this study that Sheikh Jamal Inani
National Park supports the fourth largest population of Western Hoolock Gibbons
in Bangladesh, after the larger populations in Lawachara
National Park, Adampur Reserved Forest, and Kaptai National Park (Islam et al. 2006). Ahsan (2001)
reported that the mean group size of Western Hoolock Gibbons was 3.0 (n= 8) at
West Bhanugach in northeastern
Bangladesh, whereas Feeroz & Islam (1992)
estimated a mean group size of 3.17 (n= 6) in the same area. Comparison between
group sizes in Inani and other habitats suggest a
lower reproductive output at Inani. Loss of adequate
food sources and changes in the habitat structures have led to low encounter
rates and small group sizes at the fragmented Western Hoolock Gibbon habitats
in eastern Assam, India (Kakati et al. 2009). Low
population densities have also been reported among primates in Mexico and
Brazil due to reduced food resources and habitat fragmentation (Estrada &
Coates-Estrada 1996; Chiarello & Melo 2000).
Gibbon habitat in Inani is highly degraded and
fragmented without upper canopy trees, which is likely the main reason for
their low reproductive output. We suggest that an extensive habitat restoration
programme (Hossain et al. 2008) and the total protection of gibbon habitats at Inani are required to ensure the survival of the gibbons in
this area.
Many
globally threatened wildlife species, including the Western Hoolock Gibbon, are
now on the verge of extinction at Inani due to
sharply increased pressure on natural forest resources due to the recent influx
of large numbers of Rohingya refugees from Myanmar into the area. The total
number of registered Rohingya refugees in Cox’s Bazar district is 866,457,
according to the Bangladesh Government and UNHCR, of which 716,915 are new
arrivals since 25 August 2017 (https://data2.unhcr.org/en/situations/myanmar_refugees).
Refugees
have temporarily settled in the area by clearing forests on both sides of the
Cox’s Bazar-Teknaf highway, mostly residing in the
fringes of Ukhia Reserved Forest, Inani
and Teknaf Wildlife Sanctuary, which is increasing human-wildlife
conflict in the area (Irfanullah 2018). About 3,713
acres of forest land were completely cleared to make Rohingya settlements in Ukhia, Whykheong, and Teknaf forest ranges in 2017 (UNDP Bangladesh and UN WOMEN
Bangladesh 2018). According to the Bangladesh Forest Department, an additional
6,163 acres of forest land was damaged in the areas affected by Rohingya
settlement, with no up-to-date information on the habitat status (ADB 2019).
Deforestation
and forest fragmentation, changes in forest cover, biomass reduction, loss of
species, loss of wildlife habitat, shrinkage of wildlife corridors and
increased mortality risk for wildlife are expected to result from the large
influx of migrants into Inani (UNDP Bangladesh and UN
WOMEN, Bangladesh 2018). The area influenced by Rohingya refugees is estimated
to cover 44% of the 60,000 ha landscape encompassing Sheikh Jamal Inani National Park, Ukhia Forest
Range and Teknaf Wildlife Sanctuary (UNDP Bangladesh
and UN WOMEN Bangladesh 2018), putting enormous pressure on this landscape and
the remaining forests. For example, an estimated 6,800 tons of fuel wood is
required each month by the refugee population, of which approximately 50% is
collected from the forests (UNDP Bangladesh and UN WOMEN Bangladesh 2018).
Fortunately, the Rohingya community does not hunt the gibbons. Liquefied
petroleum gas (LPG) and improved cooking stoves have been distributed since
August 2018 to Rohingya refugees and host communities to reduce the demand for
firewood from the nearby forest (IUCN Bangladesh 2019). Firewood demand dropped
by 79 % among the Rohingya families after the LPG was provided (IUCN Bangladesh
2019), but small-scale fuel-wood collection will continue to pose huge pressure
on natural resources at and around Inani.
RECOMMENDATIONS
Western
Hoolock Gibbons are likely to disappear from Sheikh Jamal Inani
National Park in the near future, if the current trend of habitat destruction
continues. The presence of large Rohingya refugee settlements have created a
critical situation that puts pressure on threatened species. Management and
conservation by the Bangladesh Forest Department of the whole of Inani is not possible due to socio-political issues and the
pressure being placed on natural resources by people living around the forest.
The Forest Department also has a shortage of manpower and other resources to
protect the large forest area. Nonetheless, the following steps can be
considered for protection and management of the gibbon habitats of Inani:
Community
members are urged to take immediate action to demarcate one designated area of
about 2,000 ha in the core gibbon habitat of Narikella
Jhuri-Bairuntali (21.229074N, 92.070104E) as a
totally protected zone. Regular monitoring and patrolling of this zone should
be prioritized by the Bangladesh Forest Department as extensive monitoring and
patrolling to the whole Inani area is not possible;
Any resource
harvesting from this core areas should be strictly prohibited and wide public
awareness campaigns must be organized to develop a positive response among the
forest-dependent people, especially fuel and timber wood collectors;
Regular
patrolling and habitat monitoring by the Forest Department should be conducted
in partnership with community patrol groups, comprising community members and
local leaders, to create a sense of stewardship and enhance protection of
forests as well as wildlife;
Highly
degraded areas identified by the Forest Department should be rehabilitated and
enriched by extensive habitat restoration programmes with native tree species,
including important food items for gibbons;
Alternative
and long-lasting poles for betel vineyards should be provided by NGOs and the
Government of Bangladesh at reasonable prices to prevent over-harvesting of
tree saplings from the forest. Extensive awareness programmes should be
conducted to discourage the collection of forest wood for poles;
Proper use
of alternative sources of fuel wood for refugees should be ensured and
regularly provided by concerned authorities; and
General
public education and awareness programs for different stakeholders should be
implemented to help to manage the globally threatened wildlife habitat of
Bangladesh on a larger scale.
Table 1. Group sizes and
composition of Western Hoolock Gibbons at Shekih
Jamal Inani National Park, Bangladesh in January
2021.
Forest jurisdiction |
Area |
Group number |
Group composition |
Total individuals |
|||||
|
|
|
AM |
AF |
SaM |
SaF |
Ju |
In |
|
Inani Range |
Inani Beat |
1 |
1 |
1 |
- |
- |
- |
- |
2 |
Inani Range |
Inani Beat |
2 |
1 |
1 |
- |
- |
- |
- |
2 |
Inani Range |
Inani Beat |
3 |
1 |
1 |
1 |
- |
- |
1 |
4 |
Inani Range |
Inani Beat |
4 |
1 |
1 |
|
1 |
1 |
- |
4 |
Inani Range |
Inani Beat |
5 |
1 |
1 |
- |
- |
- |
- |
2 |
Inani Range |
Inani Beat |
6 |
1 |
1 |
- |
- |
- |
- |
2 |
Inani Range |
Swankhali Beat |
7 |
1 |
1 |
- |
- |
- |
- |
2 |
Total |
7 |
7 |
1 |
1 |
1 |
1 |
18 |
*AM—Adult male | AF—Adult female
| SaM—Sub-adult male | SaF—Sub-adult
female | Ju—Juvenile | In—Infant.
For
images & figure - - click here
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