Journal of Threatened Taxa |
www.threatenedtaxa.org | 26 June 2020 | 12(9): 16183–16186
ISSN 0974-7907 (Online) | ISSN 0974-7893
(Print)
doi: https://doi.org/10.11609/jott.5322.12.9.16183-16186
#5322 | Received 10 August 2019 | Final
received 05 May 2020 | Finally accepted 12 May 2020
A record of gynandromorphism
in the libellulid dragonfly Crocothemis
servilia (Insecta:
Odonata) from India
R.V. Renjith
1 & A. Vivek
Chandran 2
1 Sreekailasam House, Palappuram
P.O., Ottapalam, Palakkad, Kerala 679103, India.
2 Department of Geology and
Environmental Science, Christ College, Irinjalakuda,
Thrissur, Kerala 680125, India.
1 renjith14041991@gmail.com, 2
avivekchandran@gmail.com (corresponding author)
Editor: Raymond J. Andrew, Hislop
College, Nagpur, India. Date of publication:
26 June 2020 (online & print)
Citation: Renjith,
R.V. & A.V. Chandran (2020). A record of gynandromorphism in the libellulid
dragonfly Crocothemis servilia
(Insecta: Odonata) from India. Journal of Threatened Taxa 12(9): 16183–16186. https://doi.org/10.11609/jott.5322.12.9.16183-16186
Copyright: © Renjith
& Chandran 2020. Creative Commons Attribution
4.0 International License. JoTT allows unrestricted use, reproduction, and distribution
of this article in any medium by providing adequate credit to the author(s) and
the source of publication.
Funding: None.
Competing interests: The authors
declare no competing interests.
Acknowledgements: We thank Jeevan Jose and Noppadon Makbun for helping us
confirm the condition as gynandromorphism. We are grateful to the Society for Odonate Studies, Kerala for the constant support and
encouragement. We are thankful to the
organisers and participants of Kole Odonata Survey
2019.
Gynandromorphs are genetically
and phenotypically chimeric specimens and differ from intersexes which are
genetically uniform (Narita et al. 2010).
Gynandromorphism can be bilateral, appearing
to divide down the middle into male and female sides, or they may be a mosaic,
with patches characteristic of one sex appearing in a body part characteristic
of the other sex. Gynandromorphism
is a rare phenomenon in nature and is usually detected in species that show
distinct sexual dimorphism. It is known
to occur in different arthropod taxa such as Crustacea (Farmer 2004),
Arachnida, e.g., scorpions (Cokendolpher & Sissom 1988), spiders (Palmgren
1979), ticks (Labruna et al. 2002), and Insecta, e.g., stoneflies (Klotzek
1971), hymenopterans (Gjershaug et al. 2016), beetles
(Le Gall 2006), butterflies (Nielsen 2010), dipterans such as mosquitoes (Kronefeld et al. 2013) and fruit flies (Morgan &
Bridges 1919). In vertebrates it has
been detected in reptiles (Krohmer 1989), birds (Peer
& Motz 2014), and mammals (Hollander et al.
1956). Gynandromorphs occasionally
afford a powerful tool in genetic, developmental, and behavioural
analyses. In Drosophila melanogaster,
for instance, gynandromorphs were used to provide evidence that male courtship
behaviour originates in the brain (Hotta & Benzer 1972).
Gynandromorphism is rare in odonates
(Corbet 1962) and so far has been reported from 30 individuals belonging to
seven families: Calopterygidae, Coenagrionidae,
Aeshnidae, Gomphi-dae, Cordulegastridae, Corduliidae,
and Libellulidae (Tennessen
2008; Torralba-Burrial & Ocharan
2009; Pix 2011; Futahashi
2017). There are other forms of colour
variation seen in odonates. Andromorphic females are common in many odonate families.
They have normal female reproductive organs and are fertile (Robertson
1985; Andrew 2013). Immature male
imagoes of many Anisoptera resemble females in
colouration, but can be distinguished by the male reproductive structures. Gynandromorphism,
on the contrary, is believed to be a genetic aberration caused by abnormal
mitosis in the embryo leading to unequal distribution of sex chromosomes (May
1988). As a result, the developed
individual has both male and female tissues and mixed morphological
characteristics.
The Kole
wetlands are low-lying areas that remain submerged under floodwater for about
six months of a year. Wetland agriculture, mainly paddy (rice) cultivation is
the most important activity undertaken there.
They cover an area of 13,632ha and are spread across Thrissur and
Malappuram districts of Kerala (Johnkutty &
Venugopal 1993). Kole
has been a Ramsar site since 2002, an important bird
area since 2004 (Islam & Rahmani 2008), and a
high value biodiversity area since 2009 (MoEF
2009). In a survey conducted at Kole wetlands on 14 July 2019 (Image 1), jointly by Kerala
Agricultural University, Kole Birders, Society for Odonate Studies, and Kerala Forest Department, 33 species
of odonates were recorded.
Crocothemis servilia, is a common dragonfly
associated with marshes, ponds, rivers and tanks. It is widely distributed in
the Oriental and Australian region (Subramanian 2009). The male has prominent blood red colouration
in almost all body parts including the head, thorax, abdomen and legs. The wings are transparent with the base
marked with rich amber (Image 2). The
female is pale yellow with dark brown thorax and legs (Image 3).
At Puzhakkal
region of the Kole wetlands (10.5400N
& 76.1720E), an individual of Crocothemis
servilia that looked part male and part female
was photographed during the survey. The
specimen could not be collected as it started raining and the individual moved
into the deep marshes. Subsequent
efforts to collect the specimen failed because of heavy rains that continued
for the next few days, submerging the location.
The species was initially identified using the field guide by Kiran
& Raju (2013) and confirmed by referring to Fraser (1936).
The photographed individual
showed bilateral gynandromorphism of only the thorax,
half of which showed blood red colouration as in males and the other half pale
yellow characteristic of females. The
base of the wing of the red half was marked with rich amber, in contrast with
the other wing base which was paler. The
head, legs and abdomen showed typical female morphology. Status of the anal appendages could not be
asserted from the photograph (Image 4).
Since the female characters dominated, this cannot be considered a
“balanced” gynandromorphy. The
individual exhibited a genetic mosaic condition only in the thoracic region. May (1988) reported gynandromorphism
in two species of family Corduliidae, namely Somatochlora filosa and
Somatochlora provocans
from the United States of America.
The specimens he examined had mixtures of male and female external
characters ranging from almost entirely female to about equally divided. They were symmetrical in development with
normally dimorphic structures mostly having characters intermediate between the
typical male and female conditions, particularly noticeable in the development of
the genital lobes, cerci, and metathoracic legs. Torralba-Burrial
& Ocharan (2009) reported gynandromorphism
in the libellulid Sympetrum striolatum
from Spain. One of the two specimens
they examined was a bilateral gynandromorph which looked like a female in
general appearance but had male structures in the right side of the
abdomen. Their second specimen resembled
a male in general appearance, but had a female gonopore. Mosaic gynandromorphy in Ischnura
senegalensis (Family: Coenagrionidae) and Crocothemis servilia (Family:
Libellulidae) was reported by Futasahi
from Japan (2017). In South Asia,
gynandromorphy was reported in Neurothemis tullia (Family: Libellulidae)
from Bangladesh (Shome et al. 2019), in which the
specimen’s head and thorax, including wings were bilaterally
gynandromorphic. The abdomen was androchromic but had female appendages at the tip. In India, gynandromorphism
was reported in Neurothemis tullia (Family: Libellulidae)
and Heliocypha bisignata
(Family: Chlorocyphidae) (Emiliyamma
2009), but photographs or illustrations were unavailable for confirmation or
comparison. According to Siva-Jothy (1987), gynandromorphs have been reported to fail in
mating because of aberrant sex organs but in the present individual the abdomen
is typically female and since the internal and external female reproductive
organs/genitalia are abdominal, this individual could be a fertile female. May (1988) after observing eggs on the vulvar
laminae of three Somatochlora filosa gynandromophic
individuals proposed that they were functionally female. Gynandromorphism is
a multifactorial issue caused by different genetic factors which are well
documented (Narita et al. 2010), but further research has to be undertaken to
investigate the influence of environmental factors on this phenomenon.
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