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1. Blaberidae: Rhicnoda natatrix.
a - adult male, length, 22.5mm, India, Maharashtra, Pench:
Ambakhori-Totladoah forest stream;
b - nymph, length, 10.0mm, Nepal, Makwanpur, Sim-Khola, middle
reach.
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2. Blaberidae: Rhicnoda rugosa.
a
- adult female, length 36.5mm, Nepal, Shivapuri, Dhobi-Khola;
b
- adult male length, 23.0mm, Nepal, Seti Nadi, Tributary in Pokhara Valley.
Table 1. Investigated Specimens of Rhicnoda spp. from
Nepal and India
|
Locality/Species |
Rhicnoda
natatrix |
Rhicnoda |
|
rugosa |
[body length
of |
[body length
of |
|
|
preserved |
preserved |
|
|
specimen] |
specimen] |
|
|
|
|
|
Sim-Khola,
Daman |
8.5mm nymph |
|
|
(Middle reach) |
9.5mm nymph |
|
|
|
10.0mm nymph |
|
|
|
10.5mm nymph |
|
|
|
10.5mm nymph |
|
|
First
tributary of Sim-Khola, Daman |
9.0mm nymph |
|
|
Dhobi Khola ,
Bagmati watershed |
9.5 mm nymph |
10.5mm nymph |
|
|
(not
preserved) |
36.5mm female adult |
|
Sungure Khola,
Godavari |
8.5mm nymph |
|
|
Godavari
Khola, Godavari |
10.5mm
juvenile |
11.5 nymph
(not preserved) |
|
Jakeshwori
Khola, Melamchi |
12.0mm nymph |
|
|
|
|
|
|
Pench forest
stream, spring region |
14.0mm nymph |
|
|
|
10.5mm nymph |
|
|
Pench,
Ambakori Totladoah |
13.0mm nymph |
|
|
|
18.5mm male
adult |
|
|
|
22.5mm male adult |
|
|
Forest stream,
|
|
23.0mm male
adult |
|
Seti Watershed
, Pokhara valley |
|
|
|
Total number
of specimens |
14 |
3 |
Table
2. Characteristics of different stream habitats with Rhicnoda natatrix
in Nepal and India
|
Locality |
Biocoenotic Zone, Typology |
Taxa No. |
Water Quality Class NEPBIOS |
Water Quality Class GRS-BIOS |
Water Quality Class HKHBIOS |
|
Sim
Khola midstream |
Metarhitron |
29 |
I |
I |
I |
|
First
Tributary of Sim Khola |
Epirhithron |
42 |
I |
I |
I |
|
Tributary
of Ghatte Khola* |
Epirhithron |
10 |
I |
I |
I |
|
Pench,
forest spring stream |
Hypocrenon |
12 |
I |
I |
I |
|
Ambakhori-Totladoah
forest stream* |
Metarhithron- Hyporhithron |
11 |
III |
II |
IV |
|
Average |
|
20.8 |
1.4 |
1.2 |
1.6 |
Table
3. Macro-invertebrate fauna of different Rhicnoda-habitats in Nepal
|
Abundance per Locality |
|||||||
|
Order |
Family |
I |
II |
III |
IV |
V |
|
|
Ephemeroptera |
Baetidae |
Baetis
sp. |
2 |
1 |
1 |
2 |
2 |
|
|
|
Baetiella
sp. |
1 |
|
|
|
|
|
|
|
Pseudocloeon sp. |
2 |
|
|
|
|
|
|
Heptageniidae |
|
|
|
2 |
|
1 |
|
|
|
Iron
sp. |
1 |
|
|
|
|
|
|
|
Ecdyonurus
sp. |
2 |
|
|
|
|
|
|
|
Notacanthurus
sp. |
2 |
1 |
|
|
|
|
|
Caenidae |
|
|
|
|
|
1 |
|
|
Ephemeridae
|
Ephemera
sp. |
1 |
1 |
2 |
1 |
2 |
|
|
Ephemerellidae |
|
|
|
3 |
|
3 |
|
|
Leptophlebiidae |
|
|
|
2 |
|
|
|
|
|
Indialis
sp. |
1 |
|
|
|
|
|
|
|
Habrophebiodes
sp. |
1 |
|
|
|
|
|
|
|
Serratella
sp. |
1 |
|
|
|
|
|
Plecoptera |
Perlidae |
|
|
|
4 |
3 |
|
|
|
|
Kamimura
sp. |
2 |
2 |
|
|
|
|
|
|
Tyloperla
sp. cf. |
1 |
|
|
|
|
|
|
Perlodidae |
|
|
1 |
|
|
|
|
|
Nemouridae |
|
|
|
3 |
3 |
2 |
|
|
|
Amphinemoura
sp. |
1 |
|
|
|
|
|
|
|
Mesonemoura
sp. |
|
2 |
|
|
|
|
|
Peltoperlidae |
|
2 |
|
|
|
|
|
Trichoptera |
Hydropsychidae |
Hydropsyche
sp. |
2 |
3 |
2 |
3 |
|
|
|
Odontoceridae |
|
1 |
1 |
|
|
|
|
|
Philopotamidae |
|
|
3 |
2 |
2 |
|
|
|
Limnephilidae |
Potamophylax
sp. |
|
2 |
|
|
|
|
|
Lepidostomatidae |
|
|
|
2 |
4 |
1 |
|
|
|
Paraplegopteryx
sp. |
1 |
2 |
|
|
|
|
|
Helicopsychidae |
|
|
|
2 |
|
|
|
|
Polycentropodidae |
|
|
2 |
|
|
|
|
|
Glossosomatidae |
|
|
|
|
|
1 |
|
|
Rhyacophilidae |
|
2 |
|
|
1 |
|
|
Lepidoptera |
Pyralidae |
|
|
|
1 |
|
|
|
Megaloptera |
Corydalidae |
|
|
|
1 |
|
|
|
Coleoptera |
Elmidae |
|
|
|
1 |
1 |
1 |
|
|
|
Stenelmis
sp. |
1 |
1` |
|
|
|
|
|
|
Grouvelinus
sp. |
2 |
2 |
|
|
|
|
|
Psephenidae |
|
|
|
1 |
3 |
1 |
|
|
Scirtidae |
|
2 |
2 |
1 |
2 |
1 |
|
|
Hydraenidae |
|
1 |
1 |
|
|
|
|
|
Dryopidae |
|
|
1 |
|
|
|
|
|
Dytiscidae |
|
|
1 |
2 |
1 |
|
|
|
Larvae
indet. |
|
|
|
|
1 |
|
|
Heteroptera |
Hebridae |
|
1 |
|
|
|
|
|
|
Mesoveliidae |
|
|
1 |
|
|
|
|
|
Gerridae |
|
|
|
|
|
1 |
|
Odonata |
Epiophlebiidae |
Epiophlebia
laidlawi |
2 |
2 |
|
|
|
|
|
Gomphidae |
|
|
1 |
1 |
1 |
1 |
|
|
Corduliidae |
|
|
|
|
1 |
|
|
|
Libellulidae |
|
|
|
|
|
1 |
|
|
Macromiidae |
|
|
|
|
|
1 |
|
|
Cordulegastridae |
|
|
|
1 |
1 |
1 |
|
|
Aeshnidae |
|
|
1 |
2 |
1 |
|
|
|
Euphaeidae |
|
|
|
2 |
3 |
2 |
|
|
Coenagrionidae |
|
|
1 |
|
|
|
|
|
Calopterygidae |
|
|
1 |
|
|
|
|
Diptera |
Limoniidae |
|
1 |
2 |
2 |
1 |
1 |
|
|
Athericidae |
|
|
1 |
2 |
2 |
1 |
|
|
Ceratopogonidae |
|
|
1 |
1 |
|
|
|
|
Chironomidae |
Chironominae |
|
|
|
1 |
1 |
|
|
|
Orthocladiinae |
|
2 |
1 |
|
2 |
|
|
|
Tanypodinae |
|
1 |
|
1 |
|
|
|
|
Tanytarsini |
|
2 |
|
|
|
|
|
Dolichopodidae |
|
|
1 |
|
|
1 |
|
|
Sciaridae |
|
|
2 |
|
1 |
1 |
|
|
Tabanidae |
|
|
1 |
1 |
|
|
|
|
Simuliidae |
|
1 |
2 |
|
|
3 |
|
|
Tipulidae |
|
1 |
2 |
4 |
4 |
2 |
|
|
Empididae |
|
|
|
|
1 |
1 |
|
|
Muscidae |
|
|
|
|
|
1 |
|
|
Scatophagidae |
|
|
|
|
|
1 |
|
|
Culicidae |
|
|
|
|
|
1 |
|
Blattodea |
Blaberidae |
Rhicnoda
natatrix |
1 |
2 |
1 |
1 |
|
|
|
|
Rhicnoda
rugosa |
|
|
1 |
|
1 |
|
Chilopoda |
|
|
|
1 |
|
|
|
|
Oniscoidea |
|
|
|
1 |
|
|
|
|
Decapoda |
Potamidae |
Himalayapotamon
spec. |
1 |
|
|
|
1 |
|
Tricadida |
Planariidae |
|
|
3 |
2 |
|
2 |
|
Lumbricina |
Megascolecidae |
Perionyx
fluviatilis |
1 |
1 |
2 |
|
1 |
|
Tubificina |
Tubificidae |
|
|
|
|
|
2 |
|
Hirudinida |
Salifidae |
Barbronia
nepalensis |
|
|
1 |
|
|
|
Prosobranchia |
Pomatiopsidae |
Tricula
montana |
|
|
|
2 |
1 |
|
|
|
Tricula
godavarierensis |
|
|
|
|
1 |
|
|
|
Erhaia
suguriensis |
|
|
|
2 |
|
|
Basommatophora |
Planorbiidae |
Segmentina
calatha |
|
1 |
|
|
|
|
Veneroida |
Sphaeriidae |
Pisidium
casertanum |
|
2 |
|
|
|
|
|
|
Pisidium
atkinsonianum |
|
|
2 |
|
1 |
|
|
|
Pisidium
annandalei |
|
|
|
2 |
|
|
Total
Number of Taxa |
29 |
42 |
33 |
31 |
39 |
||
Table
4. Macro-invertebrate fauna of different Rhicnoda-habitats in India
(Pench).
|
Abundance per Locality |
||||
|
Order |
Family |
Genus
species |
VI |
VII |
|
Odonata |
Libellulidae |
|
1 |
|
|
|
Platycnemididae |
|
Adults |
|
|
|
Gomphidae |
cf.
Nihonogomphus sp. |
|
2 |
|
Heteroptera |
Naucoridae |
|
1 |
1 |
|
|
Veliidae |
Microvelia sp. |
3 |
3 |
|
Coleoptera |
Dytiscidae |
|
2 |
2 |
|
|
Scirtidae |
|
1 |
|